Urumacotheriinae

PP = 60, age = 17.82 Mya (14.03–21.51).

This clade was originally proposed with fewer taxa (Negri & Ferigolo, 2004), but was expanded here to include both Urumacotherium and Pseudoprepotherium. It did not receive high node support, despite being present in trees from all analyses.

Urumacotheriinae was supported by seven synapomorphies (five for both methods and two exclusively for BI): molariform morphology of Cf1/ cf1 (BI); posterior edge of mandibular condyle inclined posterodorsally; eversion of lateral edge of symphyseal spout (BI); premolariform portion of palate roughly equal or longer than the length of molariform toothrow; presence of a distinct neck at the base of occipital condyles; occipital condyles elongated anteroposteriorly in ventral view; and lesser tubercle of humerus proximally projected as much as greater tubercle.