Published July 19, 2023 | Version v1
Taxonomic treatment Open

Stockumites convexus

  • 1. Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.

Description

Stockumites convexus (Vöhringer, 1960)

Figs 7E, 39–40; Tables 34–35

Imitoceras prorsum convexum Vöhringer, 1960: 139, pl. 2 fig. 5, text-fig. 17.

Imitoceras prorsum convexum – Weyer 1977: 172, pl. 2 figs 8–9.

Acutimitoceras convexum – Korn 1992b: 16, pl. 2 figs 17–18; 1994: 42, text-figs 49a–c, 50e, 52a, 54d, 56c. — Schönlaub et al. 1992: 16, pl. 5 figs 17–18. — Korn & Weyer 2003: 100, pl. 2 figs 1–2.

Stockumites convexus – Becker & Weyer 2004: 18, text-fig. 3g. — Korn & Weyer 2023: 30, fig. 15.

Diagnosis

Species of Stockumites with a conch reaching 40 mm diameter. Conch at 5 mm dm thickly discoidal to thinly pachyconic, subevolute to evolute (ww/dm = 0.55–0.65; uw/dm = 0.40–0.50); at 15 mm dm thickly discoidal, involute (ww/dm = 0.45–0.55; uw/dm = 0.05–0.10); at 25 mm dm thickly discoidal, involute (ww/dm = 0.45–0.55; uw/dm ~0.00). Whorl profile at 25 mm dm weakly compressed (ww/wh ~0.90); coiling rate moderate to high (WER = 1.90–2.10). Venter broadly rounded, umbilical margin broadly rounded. Growth lines coarse, wide-standing, with convex course. Weak constrictions on the shell surface; coarse internal shell thickenings. Suture line with lanceolate external lobe and V-shaped adventive lobe.

Material examined

Holotype

GERMANY • Rhenish Mountains, Oberrödinghausen railway cutting; bed 6; Vöhringer Coll.; illustrated by Vöhringer (1960: pl. 2 fig. 6) and Korn (1994: text-fig. 49a), re-illustrated here in Fig. 39A; GPIT- PV-63903.

Paratypes

GERMANY • 2 specimens; Rhenish Mountains, Oberrödinghausen, railway cutting; Hangenberg Limestone, bed 5; Vöhringer Coll.; GPIT-PV-63904, GPIT-PV-63905 • 1 specimen; Rhenish Mountains, Oberrödinghausen, railway cutting; Hangenberg Limestone, bed 6; Vöhringer Coll.; GPIT-PV-63906.

Additional material

GERMANY • 1 specimen; Rhenish Mountains, Oberrödinghausen, railway cutting; Hangenberg Limestone, bed 4; Vöhringer Coll.; MB.C.31092 • 2 specimens; Rhenish Mountains, Oberrödinghausen, railway cutting; Hangenberg Limestone, bed 5; Vöhringer Coll.; MB.C.31093.1–2 • 7 specimens; Rhenish Mountains, Oberrödinghausen, railway cutting; Hangenberg Limestone, bed 6; Vöhringer Coll.; MB.C.31094.1–7 • 1 specimen; Rhenish Mountains, Oberrödinghausen, railway cutting; Hangenberg Limestone, loose material; Korn 1977 Coll.; MB.C.31095 • 1 specimen; Rhenish Mountains, Oberrödinghausen, railway cutting; Hangenberg Limestone, bed 6b2; Weyer 1993–1994 Coll.; MB.C.31096 • 1 specimen; Rhenish Mountains, Hasselbachtal; Hangenberg Limestone, bed 59; Weyer 1993–1994 Coll.; MB.C.5241.1.

Description

Holotype GPIT-PV-63903 is a rather well-preserved specimen with 22 mm diameter; it is largely covered with shell (Fig. 39A). The conch is thickly discoidal (ww/dm = 0.51) with an almost closed umbilicus (uw/dm = 0.06) and a high coiling rate (WER = 2.08). The whorl profile is weakly compressed (ww/wh = 0.93) with convex, convergent flanks and a continuously rounded venter. The shell bears rather coarse, lamellar growth lines, which run with a broad convex arc across the flank and form a broad, distinct sinus on the venter (Fig. 40H). On the surface of the shell, there are faint constrictions that follow the course of the growth lines.

Paratype GPIT-PV-63904 (Fig. 39C) with 20 mm diameter is very similar to the holotype in its conch proportions and also the ornament. It clearly shows that the constrictions are strengthened on the inner side of the shell and lead to very deep constrictions of the internal mould. The suture line has a lanceolate external lobe with very weakly divergent flanks, a slightly wider parabolic ventrolateral saddle and a V-shaped adventive lobe with weakly outwardly curved flanks (Fig. 40G).

Smaller specimens, such as paratype GPIT-PV-63905 (12 mm dm; Fig. 39B) and especially MB.C.31096 (18.5 mm dm; Fig. 39D) show the transition from the juvenile stage to the middle stage, which is characterised by a rather rapid closure of the umbilicus by increased overlap of the inner flank area upon the umbilicus.

Specimen MB.C.31096 is well-preserved and deserves a detailed description. It has a conch diameter of 18.5 mm (Fig. 39D). It is thickly discoidal and involute (ww/dm = 0.46; uw/dm = 0.08), but shows that the umbilicus is strikingly narrowed during the last volution by strong overlap of the dorsal flank area upon the umbilicus. Half a volution before the largest diameter, the uw/dm ratio is 0.17. The umbilical margin is rounded and the flanks converge only slowly towards the broadly rounded venter; the coiling rate is high (WER = 2.08). Almost the entire specimen is covered by shell. This shows delicate growth lines, which extend with a very low and wide projection across the flank and in the ventrolateral area turn back to form a shallow ventral sinus. The shell surface bears some weak constrictions following the course of the growth lines. These constrictions begin in the inner flank area and are deepest in the ventrolateral area.

A series of cross sections demonstrates the modification of the conch from the widely umbilicate, serpenticonic juvenile stage to the discoidal subadult stage (Fig. 40 A-G). In the middle growth stage, the flanks are almost parallel; the whorl profile is widest in the middle of the convex curved flanks. The ww/dm ratio is almost constant up to a conch diameter of 30 mm (Fig. 40I). The morphological variation is particularly evident in the coiling rate. This is caused by the transformation of the serpenticonic into the disc-shaped conch at slightly different conch diameters (Fig. 40K).

Remarks

Stockumites convexus is one of the species of the genus in which the transformation of the serpenticonic juvenile stage to the involute, discoidal middle stage occurs only at a comparatively large conch diameter of about 10 mm dm. Of the species from the Rhenish Mountains, only S. antecedens is comparable in this respect, although with an even later onset of change (at 15 mm dm).

Stockumites convexus differs from the co-occurring species S. subbilobatus by the growth lines extending convexly across the flanks, which are either almost straight or slightly biconvex in S. subbilobatus.

Notes

Published as part of Korn, Dieter & Weyer, Dieter, 2023, The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany), pp. 1-230 in European Journal of Taxonomy 882 on pages 64-68, DOI: 10.5852/ejt.2023.882.2177, http://zenodo.org/record/8177581

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Linked records

Additional details

Biodiversity

Family
Prionoceratidae
Genus
Stockumites
Kingdom
Animalia
Order
Goniatitida
Phylum
Mollusca
Scientific name authorship
Vohringer
Species
convexus
Taxon rank
species
Type status
holotype , paratype
Taxonomic concept label
Stockumites convexus (Vohringer, 1960) sec. Korn & Weyer, 2023

References

  • Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196.
  • Weyer D. 1977. Ammonoideen aus dem Untertournai von Schleiz (Ostthuringisches Schiefergebirge). Zeitschrift fur geologische Wissenschaften 5: 167 - 185.
  • Korn D. 1992 b. Ammonoideen aus dem Devon / Karbon-Grenzprofil an der Grunen Schneid (Karnische Alpen, Osterreich). Jahrbuch der Geologischen Bundesanstalt 135: 7 - 19.
  • Schonlaub H. P., Attrep M., Boeckelmann K., Dreesen R., Feist R., Hahn G., Klein H. - P., Korn D., Kratz R., Magaritz M., Orth C. J. & Schramm J. - M. 1992. The Devonian / Carboniferous boundary in the Carnic Alps (Austria) - A multidisciplinary approach. Jahrbuch der Geologischen Bundesanstalt 135: 1 - 21. https: // doi. org / 10.1007 / BF 02536756
  • Korn D. & Weyer D. 2003. High resolution stratigraphy of the Devonian - Carboniferous transitional beds in the Rhenish Mountains. Mitteilungen aus dem Museum fur Naturkunde in Berlin, Geowissenschaftliche Reihe 6: 79 - 124. https: // doi. org / 10.1002 / mmng. 4860060105
  • Becker R. T. & Weyer D. 2004. Bartzschiceras n. gen. (Ammonoidea) from the Lower Tournaisian of Southern France. Mitteilungen aus dem Geologisch-Palaontologischen Institut der Universitat Hamburg 88: 11 - 36.
  • Korn D. & Weyer D. 2023. The ammonoids from the Gattendorfia Limestone of Gattendorf (Early Carboniferous, Upper Franconia, Germany). European Journal of Taxonomy 883: 1 - 61. https: // doi. org / 10.5852 / ejt. 2023.883.2179