Syzeton nubifer (LeConte, 1878), comb. nov.

(Figures 1 a, d; 2d, o; 3d; 5d)

Xylophilus nubifer LeConte, 1878: 425;

Holotype: Florida, Volusia County, Enterprise (MCZC) (Werner, 1990)

Zonantes nubifer (LeConte, 1878): Werner (1990)

Material examined. United States. Alabama: Blount County, Highland Lake, (280 m) [33°53’ N 86°25’ W], light trap, 21.VII.2013, T. King leg. (2 ♂♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 28.IV.2011, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 31. V.2014, R. Turnbow leg. (2 ♀♀, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 3.VII.2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 6.VII.2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 11.VII.2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 30.VII.2014, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 10.VIII.2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 19.VIII.2014, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 6.IV.2015, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 6.IV.2015, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 11.VI.2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 30.VI.2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 11.VII.2015, R. Turnbow leg. (6 ♀♀ and 2 ♂♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 16.VII.2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 24.VII.2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 3.VIII.2015, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 11.VIII.2015, R. Turnbow leg. (1 ♀, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 10.VI.2016, R. Turnbow leg. (1 ♂, FSCA); Dale County, Level Plains [31°17’ N 85°46’ W], black light trap, 3.VIII.2016, R. Turnbow leg. (1 ♀, FSCA). Jefferson County, Birmingham [33°31’ N 86°48’ W], black light trap, 28.IX.1981, F. N. Young leg. (1 ♂, FSCA). Florida: Alachua County, (200 m) [29°36’ N 82°22’ W], 11.IV.1992, J. Pickering leg. (2 ♀♀, FSCA); Alachua County, Highway 236, 2 km W Highway 241, 19.III.1999, C. W. Mills leg. (1 ♀, FSCA); Alachua County, Gainesville [29°39’ N 82°19’ W], black light trap, 5.IV.1983, N. Hostettler leg. (2 ♂♂, FSCA); Alachua County, Gainesville [29°39’ N 82°19’ W], black light trap,. IX.2000, electric light fixture G. Buckingham leg. (1 ♀, FSCA); A Alachua County, Gainesville, San Felasco Hammock Preserve State [29°42’ N 82°27’ W], 15.IX.1988, M. C. Thomas leg. (1 ♀, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29°37’ N 82°20’ W], 20.I.2000, P. E. Skelley leg. (1 ♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29°37’ N 82°20’ W], 4.III.2000, P. E. Skelley leg. (2 ♂♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29°37’ N 82°20’ W], live oak, black light, 10.II.2008, P. E. Skelley leg. (1 ♀, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29°37’ N 82°20’ W], black light, 24.III.2008, P. E. Skelley leg. (1 ♂, FSCA); Alachua County, Gainesville, S side Bivens Arm Lake, (30 m) [29°37’ N 82°20’ W], flight intercept trap, 27.III.2011, P. E. Skelley leg. (2 ♀♀, FSCA); Alachua County, Gainesville, 3517 NW 10th Avenue, (30 m) [29°39’ N 82°22’ W], black light trap, 24.XI.1972, R. E. Woodruff leg. (2 ♀, FSCA); Alachua County, Gainesville, 3517 NW 10th Avenue, (30 m) [29°39’ N 82°22’ W], at light, 31.VIII.2008, R. E. Woodruff leg. (1 ♀, FSCA); Alachua County, Gainesville, 610 NW 54th Terrace, (30 m) [29°39’ N 82°24’ W], 25.VIII.1994, L. Stange leg. (1 ♀, FSCA); Dade County, Camp Mahachee, South Miami, (5 m) [25°40’ N 80°16’ W], 10.IV.1993, M. C. Thomas leg. (2 ♀♀ and 1 ♂, FSCA); Escambia County, Cantonment, 2666 Monica Lane, (40 m) [30°33’ N 87°18’ W], black light trap, 10.IV.2009, T. Dobbs leg. (1 ♂, FSCA); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m), 10.IV.1991, R. Turnbow leg. (1 ♂, FSCA); Hernando County, Withlacoochee State Forest, Richloam Unit, (35 m), 27.IV.1991, R. Turnbow leg. (1 ♀ and 1 ♂, FSCA); Jackson County, Florida Caverns State Park, (20 m) [30°48’ N 85°13’ W], black light trap, 5.V.2008, M. C. & S. Thomas leg. (2 ♀♀ and 1 ♂, FSCA); Leon County, Tall Timbers Research Station, Woodyard Hammok, (60) [30°39’ N 84°12’ W], Berlese funnel, 24.XI.1972, C. W. O’Brien leg. (1 ♀, FSCA); Liberty County, Torreya State Park, (40 m) [30°33’ N 84°56’ W], black light trap, 3.V.2003, M. C. Thomas leg. (3 ♀♀, FSCA); Liberty County, Torreya State Park, (40 m) [30°33’ N 84°56’ W], 6.V.2007, M. C. Thomas leg. (2 ♀♀, FSCA); Liberty County, Aspalaga Landing, (20 m) [30°37’ N 84°54’ W], Malaise trap, 31.X.2002, DJP leg. (2 ♀♀, FSCA); Pasco County, Green Swamp West, McNeil Trailhead [28°18’ N 82°03’ W], Hardwood /coniferous forest, Beaten from branches of hardwood tree, 15.I.2014, J. P. Gruber leg. (1 ♂, WIRC); Hardwood /coniferous forest, Beaten from branches of hardwood tree, 16.I.2014, J. P. Gruber leg. (2 ♀♀, WIRC); 17.I.2014, J. P. Gruber leg. (1 ♀, WIRC); Pasco County, Green Swamp West, Main Entrance, (30 m) [28°22’02” N 82°07’04” W], Mixed hardwood/coniferous forest, Beaten from fan-like leaf of palm, 12.I.2014, J. P. Gruber leg. (1 ♀, WIRC); Pasco County, Green Swamp West, Main Entrance, (30 m) [28°22’02” N 82°07’04” W], Mixed hardwood/coniferous forest, Beaten from branches of various hardwood trees, 12.I.2014, J. P. Gruber leg. (5 ♀♀ and 1 ♂, WIRC); Pasco County, Upper Hillsborough Wildlife Management Area [28°15’32” N 82°06’50” W], Hardwood /coniferous forest, Beaten from branches of hardwood tree, 17.I.2014, J. P. Gruber leg. (1 ♂, WIRC). Georgia: Clarke County [33°54’ N 83°16’ W], 14.IX.1991, J. Pickering leg. (1 ♀, FSCA), 14.IX.1991. Louisiana: Allen Parish, Whisky Chitto Creek, (25 m) [30°41’ N 92°52’ W], Wet-mesic hardwood forest, Ex. foliage of mixed hardwoods, along streamlet; bordering Pinus cutover, 7.XI.2014, J. P. Gruber leg. (1 sp., NGPC); East Baton Rouge Parish, Baton Rouge, Place Du Plantier Apartments (5m) [30°23’55” N 91°09’13” W], Berlese forest litter, 30.XII.1993, E. G. Riley leg. (1 sp., TAMU); Rapides Parish, Indian Creek Reservoir, (25 m) [31°08’ N 92°28’ W], Dry-mesic open Pinus palustris forest, Ex. foliage of mixed hardwoods, 7.XI.2014, J. P. Gruber leg. (2 sp., NGPC). Mississippi: Oktibbeha County, Osborn Prairies, (100 m) [33°30’ N 88°44’ W], 27.V.2017, R. Turnbow leg. (1 ♀, FSCA). North Carolina: Haywood County, I-40, N of Waynesville, (840 m) [35°34’ N 82°58’ W], 2.VII.1985, H. V. Weems Jr. leg. (1 ♀, FSCA). Oklahoma: Latimer County [34°50’ N 95°18’ W], UV,. VIII.1998, K. Stephan leg. (1 sp., TAMU). South Carolina: Berkeley County, Francis Marion National Forest, Guilliard Lake Recreational Arena, (10 m) [33°17’03” N 79°37’07” W], 27.V.2006, E. G. Riley leg. (1 sp., TAMU). Texas: Brazos County, 5 km SW Wellborn, (80 m) [30°30’ N 96°20’ W], 16.VII.2016, at lights, V. Belov leg. (1 ♂, VBPC), 30.VII.2016, at lights, V. Belov leg. (1 ♂, VBPC), 23.VIII.2016, at lights, V. Belov leg. (1 ♀, VBPC), 6.IX.2016, at lights, V. Belov leg. (1 ♀, VBPC and 1 ♀, NGPC), 7.IX.2016, at lights, V. Belov leg. (1 ♀, VBPC), 8.IX.2016, at lights, V. Belov leg. (1 ♀, VBPC), 12.IX.2016, at lights, V. Belov leg. (1 ♀, VBPC), 17.IX.2016, at lights, V. Belov leg. (1 ♀, VBPC), 26.IX.2016, at lights, V. Belov leg. (1 ♂, VBPC), 27.IX.2016, at lights, V. Belov leg. (2 ♂♂, VBPC), 5.X.2016, at lights, V. Belov leg. (1 ♀, 2 ♂♂ and 1 sp., VBPC), 6.X.2016, at lights, V. Belov leg. (1 ♀, VBPC), 14.X.2016, at lights, V. Belov leg. (1 ♀, VBPC), 16.X.2016, at lights, V. Belov leg. (1 ♀, VBPC), 17.X.2016, at lights, V. Belov leg. (2 ♂♂, VBPC), 21.X.2016, at lights, V. Belov leg. (1 ♂, VBPC), 24.X.2016, at lights, V. Belov leg. (1 ♀ and 3 ♂♂, VBPC), 31. X.2016, at lights, V. Belov leg. (2 ♀♀ and 3 ♂♂, VBPC), 1.XI.2016, at lights, V. Belov leg. (3 ♂♂, VBPC), 2.XI.2016, at lights, V. Belov leg. (1 ♀ and 2 ♂♂, VBPC), 4.XI.2016, at lights, V. Belov leg. (3 ♂♂, VBPC), 18. III.2017, at lights, V. Belov leg. (1 ♀, VBPC), 27.III.2017, at lights, V. Belov leg. (1 ♀, VBPC), 29.III.2017, at lights, V. Belov leg. (1 ♂, VBPC), 13.IV.2017, at lights, V. Belov leg. (2 ♂♂, VBPC), 14.IV.2017, at lights, V. Belov leg. (1 ♀, VBPC), 15.IV.2017, at lights, V. Belov leg. (1 ♀, VBPC), 16.IV.2017, at lights, V. Belov leg. (2 ♀♀ and 1 ♂, VBPC), 18.IV.2017, at lights, V. Belov leg. (1 ♀, VBPC), 19.IV.2017, at lights, V. Belov leg. (7 ♂♂, VBPC), 21.IV.2017, at lights, V. Belov leg. (3 ♀♀ and 2 ♂♂, VBPC), 22.IV.2017, backyard, vegetation, V. Belov leg. (1 ♀ and 1 sp., VBPC), 25.IV.2017, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 1.V.2017, at lights, V. Belov leg. (1 ♀, VBPC), 13.V.2017, at lights, V. Belov leg. (2 ♀♀ and 2 ♂♂, VBPC), 15.V.2017, at lights, V. Belov leg. (1 ♂, VBPC), 19.V.2017, at lights, V. Belov leg. (1 ♀, VBPC), 4.VI.2017, at lights, V. Belov leg. (1 ♂, VBPC), 8.VI.2017, at lights, V. Belov leg. (1 ♂, VBPC), 11.VI.2017, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 20.VI.2017, at lights, V. Belov leg. (2 ♀♀ and 1 ♂, VBPC), 26.VI.2017, at lights, V. Belov leg. (1 ♂ and 2 sp., VBPC), 14.VII.2017, at lights, V. Belov leg. (1 ♂, VBPC), 8.X.2018, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 19.V.2019, at lights, V. Belov leg. (1 ♀, VBPC), 26.V.2019, at lights, V. Belov leg. (1 ♀, VBPC), 7.IV.2020, at lights, V. Belov leg. (1 ♀, VBPC), 17.IV.2020, at lights, V. Belov leg. (1 ♀, VBPC), 19.IV.2020, at lights, V. Belov leg. (1 ♂, VBPC), 20. IV.2020 at lights, V. Belov leg. (1 ♂, VBPC), 24.IV.2020, at lights, V. Belov leg. (1 ♂, VBPC), 19.V.2020, at lights, V. Belov leg. (1 ♂, VBPC), 28.V.2020, at lights, V. Belov leg. (1 ♂, VBPC), 31.V.2020, at lights, V. Belov leg. (1 ♂, VBPC), 21.VI.2020, at lights, V. Belov leg. (1 ♂, VBPC), 26.VI.2020, at lights, V. Belov leg. (1 ♂, VBPC), 3. VII.2020, at lights, V. Belov leg. (1 ♂, VBPC), 28.VII.2020, at lights, V. Belov leg. (1 ♀ and 1 ♂, VBPC), 17. IX.2020, at lights, V. Belov leg. (1 ♀, VBPC); Brazos County, College Station, Riley Estate, (80 m) [30°35’ N 96°15’ W], 27.VII.2003, E. G. Riley leg. (1 sp., TAMU), 19.IV.2005, E. G. Riley leg. (1 ♀, TAMU); Brazos County, College Station, Lick Creek Park, (70 m) [30°33’38” N 96°13’03” W], 14.IV.1996, E. G. Riley leg. (1 ♀, TAMU), 5.V.2002, E. G. Riley leg. (1 ♀, TAMU); Brazos County, Bryan [30°40’ N 96°22’ W], taken at light, 11.X.1965, J. C. Schaffner leg. (2 ♂♂, TAMU); Brazos County, Bryan [30°40’ N 96°22’ W], 13.VI.1988, E. G. Riley leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Ref., Site 8, (3 m) [25°51’06” N 97°25’13” W], Palm forest margin, Resaca bank, 20.IX.2008, E. G. Riley leg. (2 ♀♀, ERPC); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25°51’03” N 97°25’04” W], 5.IV.1987, E. G. Riley & Rider D. leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25°51’03” N 97°25’04” W], 20.X.1990, T. Carlow & E. G. Riley leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25°51’03” N 97°25’04” W], 23.IV.1994, E. G. Riley leg. (1 sp., TAMU); Cameron County, Sabal Palm Grove Sanctuary, (5 m) [25°51’03” N 97°25’04” W], 4.XI.2013, E. G. Riley & M. A. Quinn leg. (1 sp., TAMU); Fort Bend County, Brazos Bend State Park, Whiteoak Trail, (15 m) [29°22’32” N 95°34’38” W], Buckeye-sycamore forest, Beating /sweeping, 25.III.2000, E. G. Riley leg. (1 sp., TAMU); Fort Bend County, Brazos Bend State Park, Whiteoak Trail, (15 m) [29°22’32” N 95°34’38” W], Hardwood forest, Beaten from vegetation, 18.IV.2014, J. P. Gruber leg. (5 ♀♀ and 2 ♂♂, WIRC; 2 ♀♀ and 2 ♂♂, NGPC); Houston County, Crocket National Forest, Big Slough Wilderness Area, FR511 & Four C’s trail, (70 m) [31°28’ N 95°08’ W], 15.III.1996, E. G. Riley leg. (1 sp., TAMU); Montgomery County, The Woodlands [30°10’ N 95°30’ W], 25.V.1978, J. S. Wappes leg. (3 ♂♂, FSCA); Sabine County, Sabine National Forest, 9 mi. E Hemphill, Beech Bottom [31°19’ N 93°42’ W], Beech-Magnolia forest, Malaise trap, 16.IV.1989, R. Anderson & E. Morris leg. (1 sp.); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31°23’10” N 93°42’29” W], Malaise trap, 15.III.2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31°23’10” N 93°42’29” W], UV light trap, 30.III.2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31°23’10” N 93°42’29” W], Lindgren funnel, 13.IV.2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31°23’10” N 93°42’29” W], UV light trap, 10.V.2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, ca. 8 mi. E Hemphill, (50 m) [31°23’10” N 93°42’29” W], UV light trap, 15.VI.2008, E. G. Riley leg. (1 sp., TAMU); Sabine County, Sabine National Forest, 12.6 km, ESE of Hemphill [31°16’42” N 93°44’10” W], Mixed hardwood/coniferous forest, Beaten from branches of Quercus sp., 13.III.2013, J. P. Gruber leg. (2 ♂♂ and 1 sp., NGPC); Sabine County, Sabine National Forest, 12.6 km, ESE of Hemphill [31°16’42” N 93°44’10” W], Mixed hardwood/coniferous forest, Beaten from branches of Quercus sp., 14.III.2013, J. P. Gruber leg. (4♀♀ and 2 ♂♂, WIRC); San Jacinto County, Sam Houston National Forest, Big Creek Scenic Area [30°30’22” N 95°05’20” W], Mixed hardwood/coniferous forest, Beaten from branches of hardwood tree (Quercus and Carya), 11.III.2013, J. P. Gruber leg. (1 ♀ and 4 ♂♂, WIRC; 8 sp., NGPC); Tyler County, 5 mi. E Spurger, (20 m) [30°41’ N 94°05’ W], Flight intercept trap under pines, 23.III.1997, Gillogly A. R. leg. (1 sp., TAMU); Tyler County, Kirby State Forest, (40 m) [30°34’ N 94°25’ W], Elevated flight intercept, 27.IV.2003, E. G. Riley leg. (1 sp., TAMU); Tyler County, Kirby State Forest, (40 m) [30°34’ N 94°25’ W], Ground-level flight intercept trap, 27.IV.2003, E. G. Riley leg. (3 sp., TAMU); Walker County, 4 mi. N Huntsville [30°43’ N 95°33’ W], at light, 30.IV.1974, R. Phelps leg. (1 ♀, TAMU); Waller County, 15 mi. ENE of Hempstead, (80 m) [30°10’ N 95°50’ W], Beating, 22.IV.1997, Chamberlain W. E. leg. (1 sp., TAMU).

Distribution. United States: Florida to eastern Oklahoma and Texas, north to North Carolina (Werner, 1990).