Tribochrysa inaequalis Scudder 1885
- 1. Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia
- 2. Department of Earth Sciences, University of Oxford, Oxford OX 1 3 AN, United Kingdom
- 3. Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada sba 48 @ sfu. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada
Description
Tribochrysa inaequalis Scudder, 1885
Fig. 23
Tribochrysa inaequalis Scudder, 1885: 777, Fig. 982; Scudder 1890: 169, 170; Scudder 1891: 380; Handlirsch 1906–1908 [1907]: 909; Cockerell 1908a: 90; Cockerell 1909a: 78; Carpenter 1935: 267, Fig. 6; Adams 1967: 232; Schlüter 1982: 260, 261; Schlüter 1984: 4; Martins-Neto & Vulcano 1989: 189, 190, Fig. 1D; Séméria & Nel 1990: 30; Carpenter 1992: 348, Fig. 193,3; Meyer 2003: 208; Nel et al. 2005: 67; Makarkin & Archibald 2013: 143; Archibald & Makarkin 2015: 361.
Type material. Holotype MCZ 243 (Scudder’s no. 7982). A poorly-preserved articulate specimen in probably dorso-lateral aspect. Verbatim label data [typed]: (1) “Mus. Comparative Zoölogy, Cambridge / No. 243/ Tribochrysa inaequalis / Scud / Florissant Miocene”; (2) [handwritten]: “ 243 type Tribochrysa inaequalis Scud / Florissant / Neuroptera: Chrysopidae ”. The specimen has red and black ink writing on the rock that says “7982 / 243”, and a red paper label glued to the rock that says “Type.”. The specimen is covered with Canada balsam.
Type locality and horizon. USA: Colorado: Teller County: Florissant (precise collecting locality unknown); Eocene, late Priabonian; Florissant Formation.
Diagnosis. May be distinguished from T. firmata by more closely spaced subcostal veinlets (seven between origin of RP and third ra-rp) [T. firmata: five– six]; 10 branches of RP [T. firmata: eight].
Redescription. Body very poorly preserved, with only one eye well discernible (ca. 0.5 mm in diameter).
Forewing ca. 12.5 mm long, ca. 4.7 wide (length to width ratio 2.66). Costal space relatively narrow. Preserved subcostal veinlets simple, widely spaced. Pterostigma, distal part of Sc, RA not discernible. RA with nine preserved crossveins (apparently ten complete). Origin of RP relatively far from wing base (at ca. 0.22 complete length). Anterior trace of RP zigzagged, with ten branches. Basal crossvein between RP, M (1r-m) rather long, connecting anterior trace of RP, MA within im at nearly proximal one-fourth length. M arched proximad fork; dividing to MA, MP distad level of origin of RP. MA, MP strongly curved; MA rather shallowly forked distad Psc; MP deeply forked at Psc. Crossvein between MA, MP (1im) rather long. Im elongate, almost not divergent distally (length/width 4.2). Psm poorly developed, strongly zigzagged. Crossvein 2m-cu very long, connecting im, CuA at nearly proximal one-ninths im length. CuA probably with three simple branches. Psc rather well developed, slightly zigzagged. CuP deeply forked. Crossvein 2icu connecting CuA, anterior branch of CuP proximad 2m-cu. CuA, A1 closely approaching for short distance. A1, A2 simple. Two gradate series of crossveins: inner series arranged in strongly broken line, with eight preserved crossveins distad MA (apparently 8–9 complete); outer series arranged in slightly curved line, with seven crossveins distad RP2 (apparently eight complete).
Hind wing very poorly preserved.
Remarks. Much of the fossil is obscured by a covering of Canada balsam, and so we are only able to describe the forewing venation.
Notes
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Linked records
Additional details
Identifiers
Biodiversity
- Family
- Chrysopidae
- Genus
- Tribochrysa
- Kingdom
- Animalia
- Order
- Neuroptera
- Phylum
- Arthropoda
- Scientific name authorship
- Scudder
- Species
- inaequalis
- Taxon rank
- species
- Type status
- holotype
- Taxonomic concept label
- Tribochrysa inaequalis Scudder, 1885 sec. Makarkin, Antell & Archibald, 2022
References
- Scudder, S. H. (1885) Systematische Ubersicht der fossilen Myriopoden, Arachnoideen und Insekten. In: Zittel, K. A., Handbuch der Palaeontologie. Abt. 1. Palaeozoologie. Bd 2. Mollusca und Arthropoda. Verlag von R. Oldenbourg, Munchen, Leipzig, pp. 721 - 831. https: // doi. org / 10.5962 / bhl. title. 2492
- Scudder, S. H. (1890) The Tertiary insects of North America. Report of the United States Geological Survey of the Territories, 13, 1 - 734, 28 pls. https: // doi. org / 10.5962 / bhl. title. 44698
- Scudder, S. H. (1891) Index to the known fossil insects of the world including myriapods and arachnids. Bulletin of the United States Geological Survey, 71, 1 - 744. https: // doi. org / 10.5962 / bhl. title. 34455
- Cockerell, T. D. A. (1908 a) Fossil Chrysopidae. The Canadian Entomologist, 40, 90 - 91. https: // doi. org / 10.4039 / Ent 4090 - 3
- Cockerell, T. D. A. (1909 a) A catalogue of the generic names based on American insects and arachnids from the tertiary rocks, with indications of the type species. Bulletin of the American Museum of Natural History, 26, 77 - 86.
- Carpenter, F. M. (1935) Tertiary insects of the family Chrysopidae. Journal of Paleontology, 9, 259 - 271.
- Adams, P. A. (1967) A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). Bulletin of the Museum of Comparative Zoology, 135, 215 - 238.
- Schluter, T. (1982) Cimbrochrysa moleriensis n. g. n. sp. und Hypochrysa hercyniensis n. sp., zwei fossile Chrysopidae-Arten (Insecta: Planipennia) aus dem europaischen Tertiar. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 5, 257 - 264. https: // doi. org / 10.1127 / njgpm / 1982 / 1982 / 257
- Martins-Neto, R. G. & Vulcano, M. A. (1989) Neuropteros (Insecta: Planipennia) da Formacao Santana (Cretaceo Inferior), Bacia do Araripe, Nordeste do Brasil. I - Familia Chrysopidae. Anais da Academia Brasileira de Ciencias, 60, 189 - 201. [for 1988]
- Semeria, Y. & Nel, A. (1990) Paleochrysopa monteilsensis gen. et sp. nov., a new fossil of Chrysopidae from the Upper Eocene Formation of Monteils (France), with a review of the known chrysopid fossils (Insecta: Neuroptera). In: Mansell, M. W. & Aspock, H. (Eds.), Advances in Neuropterology. Proceedings of the Third International Symposium on Neuropterology (Berg in Dal, Kruger National Park, 1988). South African Department of Agricultural Development, Pretoria, pp. 27 - 32.
- Carpenter, F. M. (1992) Superclass Hexapoda. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vols. 3 & 4. Geological Society of America, Boulder, Colorado & University of Kansas, Lawrence, Kansas, pp. 1 - 655.
- Meyer, H. W. (2003) The Fossils of Florissant. Smithsonian Books, Washington, 258 pp.
- Nel, A., Delclos, X. & Hutin, A. (2005) Mesozoic chrysopid-like Planippenia: a phylogenetic approach (Insecta: Neuroptera). Annales de la Societe Entomologique de France, 41, 29 - 68. https: // doi. org / 10.1080 / 00379271.2005.10697440
- Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1
- Archibald, S. B. & Makarkin, V. N. (2015) A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the Early Eocene of Driftwood Canyon, British Columbia, Canada. Canadian Entomologist, 147, 359 - 369. https: // doi. org / 10.4039 / tce. 2014.53