Cladocalon gen. nov.

Type species: Calocladon chiriquense Gorham, 1884

Diagnosis. Cladocalon gen. nov. can be distinguished from other Eurrhacini by a combination of the following features: eyes medium to large (small in Haplobothris), pronotum as long as wide and almost parallel-sided (characteristically longer than wide and posteriorly widened in Calocladon), median longitudinal areola fusiform, present at basal 3/4 of pronotum (almost lenticular with apical half widened and present at basal 2/ 3 in Calocladon), each elytron with nine longitudinal costae (four costae in Haplobothris), abdominal sternum 9 slender and elongate (shorter and wider in Calocladon), male genitalia significantly different from all other described Eurrhacini and Calopterini: phallus with a series of folding structures, parameres flattened, slight tortuous ribbon-shaped with hook-like and/or pointed projections, widely distant in ventral view, L–shaped in lateral view (Figs. 6–15).

Description. Eyes big to medium-sized, eye diameter 1.3–1.9× longer than interocular distance. Antennomere 1 stout, 2 small, 4 about 1.2–1.5× longer than 3 (Fig. 1), antennomeres 4–10 of almost equal length, 11 twice longer than 4–10. Male antennae pectinate, reaching elytral midlength, branches of antennomeres 3–5 1.6–3× longer than antennomere length. Mandibles stout, arcuate to hooked. Labial palpi short, 3-segmented, terminal palpomere rounded. Maxillary palpi elongate, 4-segmented, palpomere 2 elongate, 4 rounded or constricted distally (Fig. 2). Labrum with distal median emargination. Anterior thoracic spiracles short. Pronotum trapezoidal (Figs. 3–4), almost as wide as long medially, almost parallel-sided, with anterior angles rounded, posterior angles produced posteriorly. Pronotum presenting median longitudinal carina with fusiform areola at basal 3/4, posterior margin of pronotum bisinuate. Scutellum rectangular, presenting apical median emargination. Elytra elongate, 4.1–4.9× longer than humeral width. Each elytron with 9 longitudinal costae, primary costae stronger, costae 2 and 4 more elevated, with mostly irregular small subsquare cells. Prosternum trapezoidal, mesosternum transverse, trapezoidal. Male abdominal sternum 8 with shallow median emargination distally, sternum 9 slender, elongate and pointed proximally, rounded distally. Trochanters elongate, conical, as long as 1/3–1/4 of femur length, tibiae as long as femur. Male genitalia slightly asymmetrical, with phallus elongate, constricted at 2/3 or at 3/4, broad medially being narrower than phallobase or as wide as phallobase in ventral view, with a series of folding structures, rounded to pointed apically, constricted at apical 1/3, basal third straight, attached to parameres proximally. Each paramere L-shaped in lateral view, flattened, slight tortuous ribbon-shaped with median/subapical/apical hook-like and/or pointed projections, widely distant in ventral view (Figs. 6–15). Phallobase in form of deep spoon. Female antennae serrate. Body length: 5.5–7.0 mm, humeral width: 1.2–1.7 mm.

Etymology: the name of the new genus is an anagram of Calocladon Gorham, 1881.

Distribution: Mexico, Guatemala and Panama.

Composition: The genus described here comprises five species: Cladocalon chiriquense comb. nov., C. histrionicum comb. nov., C. bugabense sp. nov., C. mexicanus sp. nov., and C. guatemalense sp. nov.

Comments: We found that none of syntypes of Calocladon histrionicum and C. chiriquense belong to Calocladon. In this way, two new genera should be raised to accommodate these specimens, one of them described above. The other lineage will be treated in a forthcoming article.