Apiocera Westwood, 1835

Apiocera Westwood, 1835: 448. Type species: Apiocera fiıscicollis (= A. brevicornis (Wiedemann)), by subsequent designation of Coquillett (1910: 508), Australia.

Tapinocera Macquart, 1838: 78. Type species: Laphria brevicornis Wiedemann, by monotypy, Australia.

Pomacera Macquart, 1847: 63. Type species: P. bigotii Macquart, by monotypy, Australia.

Anypenus Phillippi, 1865: 702. Type species: A. brevicornis Phillipi, by subsequent designation of Brèthes (1924), South America.

Ripidosyrma Hermann, 1909: 104. Type species: Apiocera braunsi Melander, by original designation,South Africa.

The South African species are diagnosed as follows: Head: Both sexes with eyes widely separated, frons almost parallel-sided; antennae short, flagellum two-segmented, first segment pear-shaped, second small and cylindrical with acute apex; proboscis short, hidden in buccal cavity; palpi two-segmented, second segment dorsoventrally flattened, broadly crescent-shaped. Thorax: Wings with membrane hyaline (Fig. 4), veins Ml and R5 curving forward to join wing margin before apex; discal and m3 cells present; M3 and CuA] united before wing margin, CuA2 and A1 united just before margin. Abdomen: Elongate, cigar-shaped, tapered towards apex. Male genitalia: Comparatively large; epandrium divided medially into two sclerites (Figs 6, 9); hypandrium large, independent of gonocoxites (Fig. 5); gonocoxites separate, not fused in midline; gonostyli articulated on inner margin of each gonocoxite, adjacent to base of phallus (Fig. 5); inner faces of gonocoxites with complex lobes flanking gonostyli; ejaculatory apodeme large, rounded in lateral view; lateral aedeagal apodemes relatively small, directed dorsally and posteriorly (Fig. 5). Female genitalia: Acanthophorites with row of stout spines, fused to tergite 9; three poorly sclerotised spermathecae.