Goera curvispina Martynov 1935

Goera curvispina Martynov 1935

Fig. 5

Goera curvispina Martynov 1935, 367–370, male; Akagi 1975, 3–4, larva, case; Tani 1977, 205, male; Arefina 1997, 127–128, male; Choe et al. 1999, 46, male.

Goera gyotokui Kobayashi 1957, 276–278, male. Syn. nov.

Adult. General morphology, color and size are similar to G. japonica except for the following characters. Distal segment of male maxillary palpi long oval; membranous and elastic, extended apical lobe short. In male, abdominal sternite VI with 9–15 long spines in comb-like arrangement, sternite V without spines. Female abdominal sternite VI bearing several tiny spines.

Male genitalia. Segment IX oblique, rounded anteriorly in lateral aspect. Dorsal process of tergum X absent. Paired ventrolateral processes of tergum X very long, strongly sclerotized, each tapering to acute point, and overlapped in dorsal aspect. Preanal appendages long, slender, ca. 2/3 as long as ventrolateral process. Basal segment of inferior appendage large, oval, oblique. Distal segment of inferior appendage with dorsal process strongly sclerotized, curved inward in dorsal aspect; inner mesal process strongly sclerotized, acute and curved upward in lateral aspect. Phallus slender scoop-like, membranous apically; dorsum of aedeagus with a pair of acute processes at 1/4 apically and with a pair of several thin setae at base.

Female genitalia. Preanal appendages fused with tergum X; each apex acute in dorsal aspect, concaved in lateral aspect. Supragenital plate long, relatively acute in ventral aspect. Lamellae rounded. Gonopod plate about as long as wide, apicomesal process trapezoidal. Spermathecal sclerite rectangular bearing slender anterior part in ventral aspect.

Larva. Akagi (1975) described the larval stage of this species for the first time.

Specimens examined. 1 male, holotype of G. gyotokui Kobayashi, Yoshii-machi, Ukiha-shi, Fukuoka, 17.x.1956, N. Gyotoku (MK). Hyogo: 1male, 1 female, Yasumurogawa, Takayama, Kamigori-cho, 29.x.2000, T. Murakami (TN); Shimane: 1 male, Tsuwano-cho, 17.v.1993, N. Kuhara (NK); Fukuoka: 1 male, Yoshii-machi, Ukiha-shi, 16.x.1956, N. Gyotoku (CBM); 1 female, Kanagawa, Yoshii-machi, Ukiha-machi, 17.x.1956, N. Gyotoku (TN); 1 male, Shinmachi, Chikuzen-machi, 4.iv.1972, N. Gyotoku (TN); 1 male, 1 female, Terauchi-dam, Asakura-shi, 24.iv.1993, K. Hirahara (MA). RUSSIA: Primorye: 4 males, 4 females, Ussuri River, near Zabaikalskoye village, 23.vii.1996, T. I. Arefina (TN).

Distribution. Japan (central to western Honshu, Kyushu, Tsushima), Korea, Russia (Khabarovsk, Primorye).

Japanese name. Kurubisupina-ningyo-tobikera.

Remarks. This species was described by Martynov (1935) based on material collected from south Primorye (Ussuri R.), and recorded from Japan (Lake Biwa, central Honshu) by Tsuda and Akagi (1955) for the first time. Akagi (1975) associated the larval stage with the adult based on material collected from western Honshu and Kyushu (Yamaguchi, Fukuoka and Oita). We examined males and females of this species collected from Ussuri River provided by T. I. Arefina, and determined that the Japanese specimens belong to the same species. Kobayashi (1957) described a new species, G. gyotokui, from Fukuoka, Kyushu. We examined the holotype of G. gyotokui deposited in his collection, and determined it was identical to G. curvispina. Therefore, we consider G. gyotokui to be a junior subjective synonym of G. c u r v i s p i n a. Most of Kobayashi’s collection is now preserved in CBM (Kuranishi 2002), but the type of this species has not yet moved to CBM (Kuranishi personal communication).