Archaeoditomotarsus crassitylus Faúndez, Carvajal & Rider, 2014, sp. nov.

Archaeoditomotarsus crassitylus sp. nov.

Description. Type material. Holotype. Chile, Los Lagos Region, Osorno, Puyehue, Anticura, 24-VIII-1983, leg. L. E. Peña, 1 ♂ (USNM). Paratypes: Chile, Los Lagos Region, Osorno, Puyehue, Anticura, 26, 31-VIII- 1983, leg. L. E. Peña, 1 ♂ (USNM); Chile, Los Lagos Region, Llanquihue, Chinchihuapi Creek, 41°30ʹS-73°15ʹW, 23-III- 1983, leg. T. D. Dillehay, 1 ♀ (AACN).

Head. About as long as wide; entirely covered by strong and disperse dark brown punctures which are less dense at the anterior portion (Fig. 4). First antennal segment slightly surpassing apex of clypeus, second segment shorter than third or fourth segment, third segment shorter than fourth; fifth segment lost. Eyes prominent and rounded (Fig.4). Ocelli rounded, bigger than surface punctures, not contiguous with pronotum. Posterior ending of bucculae at level with the center of the eye. Rostrum elongate, reaching or surpassing metacoxae; first rostral segment longer than the others, second segment shorter, third longer than the second, fourth segment longer than the third.

Thorax. Pronotum subtriangular, 2.5 times wider than high, humeral angles with a little cleft, slightly projected; dorsal surface covered by sparse strong brown punctures with wide unpunctate patches in the midline and towards the lateral sides. Scutellum in form of nearly equilateral triangle (Figs. 1, 2) with uniform and disperse brown punctures, and two small ivory calli on the anterolateral angles, followed by a fovea (on each side) with 1 to 4 dark brown punctures. Clavus densely punctated; hemelytra slightly convex. Corium densely punctate. Membrane hyaline with a brown patch in the middle, and few weak longitudinal veins. Thoracic sterna covered with disperse brown punctures with several unpunctate patches. Legs caramel-colored, covered by concolorous, not dense pilosoty, second tarsal segment longer than the first.

Abdomen. Connexival segments reddish-caramel in the internal half and whitish-brown in the external half, ventral portion cream-colored. Female with a dark brown patch in the middle portion of the 3 first segments of the base of the abdomen, and small dark brown near circular patches in the middle of the extant abdominal segments, male with larger patches reaching the anterior end of each abdominal segment, nearly oval-shaped; spiracles caramel-colored, situated in a concolorous or light brown patch. Female Pendergrast’s organs present on 6th and 7th sternum, forming together an ovate structure, distinctly brown colored (Fig. 5), portion on 7th sternum bigger than the portion on 6th sternum.

Female genitalia. First gonocoxae elongated and wide, with rugose aspect, each gonocoxite convex posteriorly, each sclerite subdeltoid shaped (Fig. 5); second gonocoxae small with each gonocoxite nearly rectangular posteriorly (Fig. 5); paratergites 8 rounded posteriorly; paratergites 9 acute in the posterior portion and concave anteriorly, with a little projection extending laterally (Fig. 5).

Male genitalia: Pygophore slightly punctate, ventral rim slightly concave medially (Fig. 7), which is visible in dorsal and ventral view; two small black processes are visible on posterolateral angles in dorsal view. Parameres wide and elongated, with a y-shaped process at the basal portion, oriented to the messial part of pygophore, apical portion of parameres sharpened (Fig. 8).

Measurements (mm; male followed by female in parentheses). Total length: 7.01 (7.90); head length: 1.23 (1.34), head width: 1.28 (1.38); antennal segments: I—0.61, II—0.83, III—0.91, IV—1.32, V: lost; rostral segments: I—0.90, II—0.61, III—0.74, IV—0.88; pronotum length: 1.52 (1.72), pronotum width: 3.86 (4.36); scutellum length: 2.07 (2.19), scutellum width: 2.15 (2.26).

Etymology. The species name is composed from the Latin crassi -, meaning thick, and tylus, meaning anteclypeus, in reference of the prominent anteclypeus of this species; noun in apposition.