Condyloderes shirleyi Neuhaus & Zotto & Yamasaki & Higgins 2019, sp. nov.

Description of Condyloderes shirleyi sp. nov. Neuhaus & Higgins

(Figs 33–44; Tables 1, 5–7) http://zoobank.org/ urn:lsid:zoobank.org:act: 0A136401-7882-45BA-A92C-9D1FA76E08EA

Condyloderes sp.—Neuhaus 2013: pp. 232, 239, 267, figs 5.1.14, 5.1.15.—Neuhaus 2017: p. 146.

Diagnosis. Neck placids with condyles in two rows in narrower placids and three rows in broader placids; each narrower placid with one apical and two basal condyles except paraventral placid next to midventral placid revealing in majority of specimens one apical and one basal condyle; each broader placid usually with two apical condyles, two condyles in intermediate row, and four basal condyles; midventral placid always with three apical condyles and usually two in intermediate row and four in basal row; acicular spine middorsally on segments 1–9 (and on segment 10 in male only), midterminally on segment 11, lateroventrally on segments 1–9, and laterodorsally on segment 10 (in male only); cuspidate spine ventrolaterally on segments 2, 5, 7 and 9, lateral accessorily on segments 2, 4, 6 and 8, and middorsally on segments 5 and 7; type-5 (?) sensory spot ventromedially on segment 11; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally) and 29, lateroventrally on segment 10, sublaterally on segments 1–3, 5-7 and 9, midlaterally on segments 1 and 2, laterodorsally on segments 1–9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2 and 4–9; gland cell outlet ventromedially on segment 10 in male only; ventromedial appendage on segments 6–8 and area of micropapillae ventromedially on segment 9 in female only.

Material examined. Holotypic male (USNM 1480999), 44 paratypic males (USNM 1480980-1480998, 1481000-1481019; ZMB 11598-11602), 27 paratypic females (USNM 1481020-1481029, 1481031-1481042; ZMB 11603-11607), and one paratypic female exuvia (USNM 1481030), mounted for light microscope study; 6 males and 5 females, mounted for examination with the SEM (USNM 1481164, 1481166-1481172). All specimens originate from the same sample in Alaska (Fig. 1; Table 1).

Deposition of material. The holotype and 62 paratypes are deposited at the National Museum of Natural History, Smithsonian Institution, Washington D.C., catalogued under the numbers USNM 1480999 (holotype) and USNM 1480980-1480998, 1481000-1481042 (Table 1). Also, all specimens mounted for SEM investigation are kept at the Smithsonian Institution (USNM 1481164, 1481166-1481172). Ten additional paratypes are located in the “worm” collection of the Museum für Naturkunde, Berlin and catalogued in the “Generalkatalog freilebende Würmer” under the numbers ZMB 11598-11607 (Table 1).

Type locality. North Pacific Ocean, Alaska, Glacier Bay, Berg Bay, North Element, 58°32’48’’ N, 136°09’00’’ W.

Etymology. The species is named in honour of Prof. Dr Thomas Shirley, who supported sampling in Alaska.