Biodiversity Literature Repository
Published October 6, 2009 | Version v1
Journal article Open

Proposal of Marbefia, gen. n. and Inermiphonte, gen. n., including updated keys to the species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948 (Copepoda, Harpacticoida, Laophontidae)

Creators

  • 1. Natural History Museum, London, United Kingdom
  • 2. Hanyang University, Laboratory of Biodiversity, Seoul, Korea, South

Description

Both sexes of the enigmatic laophontid species Pseudonychocamptus carthyi Hamond, 1968 (Copepoda, Harpacticoida) are redescribed in detail based on type and other material from Norfolk, England. The species exhibits marked differences with other Pseudonychocamptus species and is consequently fixed as the type of a new genus Marbefia, gen. n., being defined by the following autapomorphies: (1) P2–P4 enp-2 (except P4 enp-2 in ♂) inflated, with two parallel rows of flimsy setular extensions; (2) P4 with sexually dimorphic setation on enp-2 (outer distal seta reduced in ♀, very long in ♂); and (3) antennule ♀ with spinules along posterior margins of segments 1–6. Laophonte danversae Hamond, 1969, "Laophontid male, ?gen., ?sp." sensu Hamond (1969) [renamed as Inermiphonte hamondi, sp. n.] and Laophonte ?drachi Médioni & Soyer, 1966 are collectively placed in a new genus Inermiphonte, gen. n. which shares as sistergroup relationship with Harrietella T. Scott, 1906. Marbefia, gen. n. is considered sister to a clade comprising the genera Pseudonychocamptus Lang, 1944, Pilifera Noodt, 1952, Inermiphonte, gen. n. and Harrietella. Pseudonychocamptus kolarovi Apostolov, 2008 is transferred to Paralaophonte whereas Chislenko's (1967) record of P. koreni Sars, 1908a is considered doubtful. Willey's (1935) variety "fissirostris" of Paralaophonte brevirostris (Claus, 1863) is rejected pending a thorough analysis of the variability in the P. brevirostris complex of species. P. congenera mediterranea Lang, 1948 appears to be more closely related to P. lacerdai Jakobi, 1953 than to its nominotypical subspecies P. congenera congenera (Sars, 1908b) whereas P. pacifica galapagoensis Mielke, 1981 (here upgraded to full specific rank) is more similar to P. brevirostris than to its nominotypical subspecies P. pacifica pacifica Lang, 1965. A new species, Paralaophonte pallaresae, sp. n. is proposed for P. gracilipes Brady, 1910 sensu Pallares (1968). The potentially paraphyletic status of the genus Paralaophonte is discussed. Dichotomous identification keys are provided for the valid species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948.

Files

ZK_article_2079.pdf

Files (2.9 MB)

Name Size Download all
md5:734187651608cb5fc30c1b5844e2d275
2.9 MB Preview Download

Linked records

Additional details

Identifiers

LSID
urn:lsid:zoobank.org:pub:A447D3B6-5387-44B6-AC1B-A05988000E43
LSID
urn:lsid:plazi.org:pub:734187651608CB5FC30C1B58FFE2D275
URL
http://publication.plazi.org/id/734187651608CB5FC30C1B58FFE2D275

Has part
Taxonomic treatment: 10.5281/zenodo.3790664 (DOI)
Taxonomic treatment: urn:lsid:zoobank.org:act:5B070367-A759-44B2-931C-7D9A49AC6AC4 (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8F78FF1D160BCB5BC38018D1FC0FD621 (URL)
Taxonomic treatment: 10.5281/zenodo.3790666 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/8F78FF1D160CCB4FC3801FFBFC0ED081 (URL)
Taxonomic treatment: 10.5281/zenodo.3790668 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/8F78FF1D1618CB4EC380181BFBB1D6C8 (URL)
Taxonomic treatment: 10.5281/zenodo.3790670 (DOI)
Taxonomic treatment: urn:lsid:zoobank.org:act:EB895638-2BFC-408F-8B2C-00B93A2480DE (LSID)
Taxonomic treatment: http://treatment.plazi.org/id/8F78FF1D1619CB4BC3801E44FE24D4FD (URL)
Taxonomic treatment: 10.5281/zenodo.3790672 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/8F78FF1D161DCB43C3801BFFFBB6D4F2 (URL)
Figure: 10.5281/zenodo.3767614 (DOI)
Figure: 10.5281/zenodo.3767616 (DOI)
Figure: 10.5281/zenodo.3767618 (DOI)
Figure: 10.5281/zenodo.3767620 (DOI)
Figure: 10.5281/zenodo.3767622 (DOI)
Is source of
Dataset: http://www.gbif.org/dataset/6b003da1-30a4-4961-839d-53f2f79ae563 (URL)

References

  • Apostolov A (2004) Paralaophonte livingstoni n. sp. (Crustacea: Copepoda: Harpacticoida) - un nouveau representant de l'ile de Livingstone, Antarctique. Historia naturalis bulgarica 16: 59-67.
  • Apostolov A (2008) Harpacticoides (Crustacea, Copepoda) de la mer Egee (plages de Kavala, Grece du nord). Historia naturalis bulgarica 19: 5-33.
  • Apostolov A, Petkovski T (1980) Copepodes Harpacticoides de la mer Adriatique (region de Rovigne). Acta zoologica bulgarica 15: 24-35.
  • Bodin P (1971) Catalogue des nouveaux Copepodes Harpacticoides marins. Additif no. 1. Tethys 2(4): 881-907.
  • Bodin P (1979) Catalogue des nouveaux Copepodes Harpacticoides marins. (Nouvelle edition). Universite de Bretagne occidentale. Laboratoire d'Oceanographie biologique: 1-228.
  • Bodin P (1997) Catalogue of the new marine harpacticoid copepods (1997 edition). Documents de Travail, Institut royal des Sciences naturelles de Belgique 89: 1-304.
  • Boeck A (1873) Nye Slaegter og Arter af Saltvands-Copepoder. Forhandlinger i Videnskabsselskabet i Kristiania 1872: 35-60.
  • Brady GS (1899) On the marine Copepoda of New Zealand. Transactions of the zoological Society of London 15(2): 31-54.
  • Brady GS (1910) Die marinen Copepoden der Deutschen Sudpolar Expedition 1901-1903. I. Ueber die Copepoden der Stamme Harpacticoida, Cyclopoida, Notodelphyoida und Caligoida. Deutsche Sudpolar-Expedition 11 (= Zoology 3): 497-594.
  • Brian A (1917) Descrizione di una nuova specie di Laophonte (L. quaterspinata n. sp. mihi) proveniente dai materiali del Laboratorio marino di Quarto. Monitore zoologico Italiano 28: 174-178.
  • Brian A (1921) I Copepodi Harpacticoidi del Golfo di Genova. Stab. Tip.-Lit. R. Istituto Sordomuti, Genova, 112 pp.
  • Ceccherelli VU (1988) Trois nouvelles especes de Copepodes Harpacticoides des lagunes de l'Adriatique du Nord, Italie. Vie et Milieu 38(2): 155-171.
  • Chislenko LL (1967) Garpaktitsidy (Copepoda Harpacticoida) Karelskogo poberezh'ya Belogo morya. Copepoda Harpacticoida of the Karelian coast of the White Sea. In: Gidrobiologichyeskiye Issledovaniya na Karel'skom poberezh'e Belogo moray (Hydrobiological Researches of the Karelian coast of the White Sea). Issledovaniya Fauny Morei (Explorations of the Fauna of the Seas) 7(15): 48-196 [In Russian].
  • Chislenko LL (1977) Garpaktitsidy (Copepoda, Harpacticoida) s gubok zemli Frantsa-Iosifa. Harpacticids (Copepoda Harpacticoidea) from sponges of Franz Josef Land. In: Biotsenosy shel'fa zemli Frantsa-Iosifa i fauna sopredel'nykh akuatorii. Issledovaniya Fauny Morei (Explorations of the Fauna of the Seas) 14(22): 237-276 [In Russian].
  • Chislenko LL (1978) New species of copepod harpacticoids (Copepoda, Harpacticoida) from the Possjet Bay of the Sea of Japan. Trudy zoologicheskogo Instituta, Akademiia nauk SSSR 61: 161-192 [In Russian].
  • Claus C (1863) Die freilebenden Copepoden mit besonderer Berucksichtigung der Fauna Deutschlands, der Nordsee und des Mittelmeeres. Wilhelm Engelmann, Leipzig, 230 pp.
  • Douwe, C van, (1929) Marine Litoral-Copepoden: Zur Verbreitung des Genus Laophonte Philippi im Mittelmeer. Zoologischer Anzeiger 83: 283-294.
  • Fiers F (1986) Harpacticoid copepods from the West Indian Islands: Laophontidae (Copepoda, Harpacticoda). Amsterdam Expedition to the West Indian Islands, Report 48. Bijdragen tot de Dierkunde 56(1): 132-164.
  • Fiers F (1992) A redescription of Hemilaophonte janinae Jakubisiak (Copepoda, Harpacticoida), a laophontid living in the gill chambers of the common spider crab. Belgian Journal of Zoology 122: 211-222.
  • Fiers F (1993) The laophontid genus Loureirophonte Jakobi, 1953 (Copepoda, Harpacticoida). Zoologische Mededelingen, Leiden 67: 207-238.
  • Gheerardyn H, Fiers F, Vincx M, De Troch M (2006) Paralaophonte harpagone, sp. n. (Copepoda: Harpacticoida), a laophontid with an extremely specialised maxilliped. Organisms, Diversity and Evolution 6(3): 323-324 [Extended abstract]. [Full description in Electronic Supplement 14: 1-9; http://www.senckenberg.de/odes/06-14.htm - last accessed 07 April 2009].
  • Goddard M (2006) Copepodos de pozas intermareales de Isla San Felix y del Archipielago de Juan Fernandez. Tide pool copepods of San Felix Island and Juan Fernandez Archipelago. Ciencia y Tecnologia del Mar 29: 115-122.
  • Gomez S, Boyko CB (2006) On a small collection of harpacticoids from Easter Island: the family Laophontidae T. Scott (Crustacea: Copepoda: Harpacticoida). Zootaxa 1352: 3-70.
  • Gurney R (1927) Report on the Crustacea: -- Copepoda (littoral and semi-parasitic). Zoological results of the Cambridge expedition to the Suez Canal, 1924, no. 35. Transactions of the Zoological Society of London 22: 451-577.
  • Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172-176.
  • Hamond R (1969) The Laophontidae (Copepoda, Harpacticoida) of the shore at West Runton, Norfolk, England. Crustaceana 16: 1-14.
  • Hamond R (1972) Some marine and brackish-water copepods from Wells-next-the-Sea, Norfolk, England. Transactions of the Norfolk and Norwich Naturalists' Society 22: 237-243.
  • Hamond R (1973) Some Laophontidae (Crustacea: Harpacticoida) from off North Carolina. Transactions of the American microscopical Society 92: 44-59.
  • Hicks GRF (1980) Structure of phytal harpacticoid copepod assemblages and the influence of habitat complexity and turbidity. Journal of experimental marine Biology and Ecology 44: 157-192.
  • Hicks GRF (1982) New records of harpacticoid copepods from the east coast of Britain, and a description of the true male of Laophonte danversae Hamond. Crustaceana 42: 302-307.
  • Hicks GRF (1989) Harpacticoid copepods fom biogenic substrata in offshore waters of New Zealand. 2. Partial revisions of Dactylopodella Sars and Amphiascus Sars (varians -group) including new species, and a new record for Harrietella simulans (T. Scott). National Museum of New Zealand Records 3(10): 101-117.
  • Holmes JMC, Minchin D (2000) Pseudonychocamptus carthyi Hamond (Crustacea: Copepoda: Harpacticoida) new to Ireland. Bulletin of the Irish Biogeographical Society 23: 141-142.
  • Huys R (2009a) On the junior subjective synonyms of Coullia Hamond, 1973 (Copepoda, Harpacticoida, Laophontidae): an update and key to species and related genera. ZooKeys 5: 33-40.
  • Huys R (2009b) Unresolved cases of type fixation, synonymy and homonymy in harpacticoid copepod nomenclature (Crustacea: Copepoda). Zootaxa 2183: 1-99.
  • Huys R, Gee JM, Moore CG, Hamond R (1996) Marine and Brackish Water Harpacticoid Copepods. Part 1. In: Barnes RSK, Crothers JH (Eds), Synopses of the British Fauna (New Series) 51: i-viii, 1-352. Field Studies Council, Shrewsbury.
  • Huys R, Iliffe TM (1998) Novocriniidae, a new family of harpacticoid copepods from anchihaline caves in Belize. Zoologica Scripta 27: 1-15.
  • Huys R, Lee W (2000) Basal resolution of laophontid phylogeny and the paraphyly of Esola Edwards. Bulletin of the Natural History Museum, London (Zoology) 66: 49-107.
  • Jakobi H (1953) Novos Laophontidae (Copepoda-Crustacea) da costa Brasileira. (Neue Laophontiden von der Brasilianischen Kuste). Dusenia 4: 47-60.
  • Jakubisiak S (1932) Sur les Harpacticoides heberges par Maia squinado. Bulletin de la Societe zoologique de France 57: 506-513.
  • Klie W (1929) Die Copepoda Harpacticoida der sudlichen und westlichen Ostsee mit besonderer Berucksichtigung der Sandfauna der Kieler Bucht. Zoologische Jahrbucher fur Systematik 57(3-4): 329-386.
  • Kornev PN, Chertoprud ES (2008) Harpacticoida copepods of the White Sea: morphology, systematics, ecology. KMK Scientific Press Ltd, Moscow, 379 pp.
  • Lang K (1934) Marine Harpacticiden von der Campbell-Insel und einigen anderen sudlichen Inseln. Acta Universitatis lundensis, n. ser. Avd. 2, 30(14) : 1-56.
  • Lang K (1936) Copepoda Harpacticoida. Further zoological Results of the Swedish Antarctic Expedition 1901-1903 under the Direction of Dr. Otto Nordenskjold 3(3): 1-68.
  • Lang K (1944) Monographie der Harpacticiden (Vorlaufige Mitteilung). Almqvist & Wiksells Boktryckeri Ab, Uppsala, 39 pp.
  • Lang K (1948) Monographie der Harpacticiden. Hakan Ohlsson, Lund, 1682 pp. (2 volumes).
  • Lang K (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska Vetenskapsakademiens Handlingar (4)10(2): 1-560.
  • Lee W, Huys R (1999) Bathylaophonte, gen. n. from deep-sea hydrothermal vents and the polyphyly of Paronychocamptus (Copepoda: Harpacticoida). Cahiers de Biologie marine 40: 293-328.
  • Letova VN (1982) Harpacticoida (Crustacea, Copepoda) ilisto peschano litorali vostochnogo Murmana. Harpacticoida (Crustacea, Copepoda) from the mud-sandy littoral of the east Murman. In: Skarlato OA (ed.), Bespozvonochnye pribrezhnykh biotsenozov severnogo ledovitego (Marine invertebrates of coastal biocenoses of the Arctic Ocean and the Pacific Ocean). Issledovaniya Fauny Morei (Explorations of the Fauna of the Seas) 29(37): 46-75 [in Russian].
  • Medioni A, Soyer J (1966) Laophonte ? drachi n. sp. Copepode Harpacticoide associe au Bryozoaire Schismopora armata (Hincks, 1860). Vie et Milieu (A)17(2): 1053-1063.
  • Mielke W (1974) Eulitorale Harpacticoidea (Copepoda) von Spitzbergen. Mikrofauna des Meeresbodens 37: 1-52.
  • Mielke W (1975) Systematik der Copepoda eines Sandstrandes der Nordseeinsel Sylt. Mikrofauna des Meeresbodens 52: 1-134.
  • Mielke W (1981) Interstitielle Fauna von Galapagos. XXVIII. Laophontinae (Laophontidae), Ancorabolidae (Harpacticoida). Mikrofauna des Meeresbodens 84: 1-106.
  • Mielke W (1982) Einige Laophontidae (Copepoda, Harpacticoida) von Panama. Crustaceana 42(1): 1-11.
  • Mielke W (2001) Loureirophonte psammophila, a new species of Laophontidae T. Scott, 1904 (Copepoda: Harpacticoida) from Baja California, Mexico. Hydrobiologia 445: 77-83.
  • Monard A (1928) Les Harpacticoides marins de Banyuls. Archives de Zoologie experimentale et generale 67: 259-443.
  • Monard A (1935) Les Harpacticoides marins de la region de Salammbo. Bulletin de la Station oceanographique de Salammbo 34: 1-94.
  • Monard A (1936) Note preliminaire sur la faune des Harpacticoides marins d'Alger. Bulletin de la Station d'Aquiculture et de Peche de Castiglione 1935(1) : 45-85.
  • Nicholls AG (1944) Littoral Copepoda from the Red Sea. Annals and Magazine of natural History (11)11: 487-503.
  • Nicholls AG (1945) Marine Copepoda from Western Australia. III. - Littoral harpacticoids from Port Denison. Journal of the Royal Society of Western Australia 29: 1-16.
  • Noodt W (1952) Marine Harpacticiden (Cop.) aus dem eulitoralen Sandstrand der Insel Sylt. Abhandlungen der mathematisch-naturwissenschaftlichen Klasse. Akademie der Wissenschaften und der Literatur in Mainz 1952(3): 105-142.
  • Noodt W (1958) Die Copepoda Harpacticoidea des Brandungsstrandes von Teneriffa (Kanarische Inseln). Abhandlungen der mathematisch-naturwissenschaftlichen Klasse. Akademie der Wissenschaften und der Literatur in Mainz 1958(2): 53-116.
  • Nurul Huda AI, Zaleha K (2005) Report on some phytal harpacticoid copepods from Terengganu coast. Proceedings of the KUSTEM 4th Annual Seminar 2005. Primula Beach Resort, Kuala Terengganu, Terengganu: 392-396.
  • Pallares RE (1968) Copepodos marinos de la Ria Deseado (Santa Cruz, Argentina). Contribucion sistematica-ecologica. I. Contribucion cientifica. Centro de Investigacion de Biologia Marina, Buenos Aires 27: 1-125.
  • Pallas PS (1766) Elenchus zoophytorum sistens generum adumbrationes generaliores et specierum cognitarum succinctas descriptiones cum selectis auctorum synomymis. F. Varrentrapp, Hagae, xxviii + 451 pp.
  • Pesta O (1959) Harpacticoiden (Crust. Copepoda) aus submarinen Hohlen und den benachbarten Litoralbezirken am Kap von Sorrent (Neapel). Ergebnisse der Osterreichischen Tyrrhenia Expedition 1952. Teil: VI. Pubblicazione della Stazione zoologica di Napoli 30(suppl.): 94-177.
  • Petkovski T (1964) Zur Kenntnis der Harpacticiden Portugals (Crustacea, Copepoda). Acta universitatis lundensis, n.ser., Avd 2, 59(14): 1-22.
  • Philippi A (1840) Zoologische Bemerkungen (Fortsetzung). IV. Kurze Charakteristik mehrerer neuer Genera aus der Familie der Copepoden. Archiv fur Naturgeschichte 6: 188-190.
  • Por FD (1964) A study of Levantine and Pontic Harpacticoida (Crustacea, Copepoda). Zoologische Verhandelingen, Leiden 64: 1-128.
  • Raibaut A (1969) Paralaophonte ormieresi n. sp. Copepode HarpacticoVde trouve sur les branchies de Maia squinado (Herbst) (Crustacea, Decapoda). Bulletin de la Societe zoologique de France 93(3): 451-457.
  • Sars, GO (1903) Copepoda Harpacticoida. Parts I & II. Misophriidae, Longipediidae, Cerviniidae, Ectinosomidae (part). An account of the Crustacea of Norway, with short descriptions and figures of all the species 5: 1-28 + pls. 1-16.
  • Sars GO (1905) Pacifische Plankton-Crustaceen. (Ergebnisse einer Reise nach dem Pacific. Schauinsland 1896-1897). II. Brackwasser-Crustaceen von den Chatham-Inseln. Zoologische Jahrbucher fur Systematik 21(4): 371-414.
  • Sars GO (1908a) Copepoda Harpacticoida. Parts XXI & XXII. Laophontidae (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 5: 241-256 + pls. 161-176.
  • Sars GO (1908b) Copepoda Harpacticoida. Parts XXIII & XXIV. Laophontidae (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 5: 257-276 + pls. 177-192.
  • Sars GO (1909) Crustacea. Report of the second Norwegian Arctic Expedition in the "Fram" 1898-1902 18: 1-47.
  • Sars GO (1911) Copepoda Harpacticoida. Parts XXXV & XXXVI. Supplement (concluded), index, etc. An account of the Crustacea of Norway, with short descriptions and figures of all the species 5: 421-449 + i-xiv (text) + i-xii (plates) + title of text and of plates + supplement pls. 43-54.
  • Sars GO (1920a) Copepoda Supplement. Parts V & VI. Harpacticoida (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 7: 53-72 + pls. 33-48.
  • Sars GO (1920b) Copepoda Supplement. Parts VII & VIII. Harpacticoida (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 7: 73-92 + pls. 49-64.
  • Scott T (1894a) Additions to the fauna of the Firth of Forth. Part VI. Reports of the Fishery Board for Scotland, Edinburgh 12(3): 231-271.
  • Scott T (1894b) Report on Entomostraca from the Gulf of Guinea, collected by John Rattray, B.Sc. Transactions of the Linnean Society of London, Zoology (2)6: 1-161.
  • Scott T (1899) Report on the marine and freshwater Crustacea from Franz-Josef Land, collected by Mr. William S. Bruce, of the Jackson-Harmsworth Expedition. Journal of the Linnean Society, Zoology 27: 60-126.
  • Scott T (1903) On some new and rare Crustacea collected at various times in connection with the investigations of the Fishery Board for Scotland. Reports of the Fishery Board for Scotland, Edinburgh 21(3): 109-135.
  • Scott T (1905) On some new and rare Crustacea from the Scottish seas. Reports of the Fishery Board for Scotland, Edinburgh 23(3): 141-153.
  • Scott T (1906) Notes on British Copepoda: change of names. Annals and Magazine of Natural History 7(17): 458-466.
  • Sewell RBS (1940) Copepoda, Harpacticoida. The John Murray Expedition 1933-34 Scientific Reports 7(2): 117-382.
  • Vervoort W (1950) Harrietella simulans (T. Scott, 1894), a commensal copepod on Limnoria lignorum (Rathke). Zoologische Mededelingen Leiden 30: 297-305.
  • Vervoort W (1962) Report on some Copepoda collected during the Melanesia Expedition of the Osaka Museum of Natural History. Publications of the Seto marine biological Laboratory 10: 393-470.
  • Vervoort W (1964) Freeliving Copepoda from Ifaluk Atoll in the Caroline Islands with notes on related species. Bulletin of the United States National Museum 236: i-ix, 1-431.
  • Wells JBJ (1970) The marine flora and fauna of the Isles of Scilly. Crustacea: Copepoda: Harpacticoida. Journal of natural History 4(2): 255-268.
  • Wells JBJ (1976) Keys to aid in the identification of marine harpacticoid copepods. Published by the Department of Zoology, University of Aberdeen, U.K.: 1-215.
  • Wells JBJ (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa 1568: 1-872.
  • Wells JBJ, Hicks GRF, Coull BC (1982) Common harpacticoid copepods from New Zealand harbours and estuaries. New Zealand Journal of Zoology 9: 151-184.
  • Wells JBJ, Rao GC (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India 16: 1-385.
  • Willen E (1996) Two new genera of Laophontidae (Copepoda: Harpacticoida) from the high Antarctic Weddell Sea. Journal of natural History 30: 1297-1327.
  • Willey A (1930) Harpacticoid Copepoda from Bermuda. - Part I. Annals and Magazine of natural History (10)6: 81-114.
  • Willey A (1935) Harpacticoid Copepoda from Bermuda. - Part II. Annals and Magazine of natural History (10)15: 50-100.
  • Wilson CB (1932) The copepods of the Woods Hole region, Massachusetts. Bulletin of the United States National Museum 158: 1-635.
  • Yeatman HC (1970) Copepods from Chesapeake Bay sponges including Asterocheres jeanyeatmanae n. sp. Transactions of the American microscopical Society 89: 27-38.
  • Yoo K-I, Lee W (1995) Marine harpacticoid copepods from the Korean waters. The Yellow Sea 1: 34-49.
  • Zaleha K, Roswati MA, Iwasaki N (2006) Distribution of some species of harpacticoid copepods in east coast of Peninsular Malaysia. Coastal marine Science 30: 140-145.