Cleptoria Chamberlin 1939

Material examined: Type specimens. ♂ holotype (VTEC MPE00291), 1 ♂ paratype (VMNH SPC000389), and 2 ♀ paratypes (VTEC SPC000390, 394). Holotype from South Carolina, Barnwell County, Toby Creek, near Hagoods Mill, SC-64 (33.2119°N, - 81.3201°W, Elev: 46 m), 23 November 2003, (Coll: D. Beamer). Paratypes from South Carolina, Barnwell County, Hagoods Mill, SC-64, 351 rd m W jct w/ Lebanon Rd., nr Toby Ck (33.21206°N, - 81.3198°W, Elev: 46 m), 17 October 2004, beneath leaf litter (Quercus, Acer) (Coll: P. Marek, D. Beamer, M. Beamer). Material examined listed in Supp Table 2 (online only).

Diagnosis: Adult males of Sigmoria beameri n. sp. are distinct from other apheloriine species based on the following combination of characters: Gonopods. Gonopodal acropodite broad, curving dorsomedially at an almost 90° angle at anterior bend (Fig. 13A). Basal zone lacking spines or tubercles, prefemoral process absent. Medial flange thick, laminate. Acropodite peak broad, flattened, distally with a wide, laminate lateral flange which is distinctly constricted at its base, setting it off from its ventrally-directed tip, separating it from other species with a birdhead-like acropodal apex (= Cleptoria) Sigmoria abbotti (Hoffman, 1967), Sigmoria bipraesidens (Hoffman, 1967), Sigmoria macra (Chamberlin, 1939), Sigmoria rileyi (Bollman, 1889), Sigmoria robusta Shelley, 1986, Sigmoria shelfordi (Loomis, 1944), and Sigmoria arcuata (Shelley, 1981). Acropodite without a cingulum, as in Brachoria and Appalachioria. Color. Tergites with light-red paranotal spots (Fig. 5R). Dark brown background. Collum with light red anterior and lateral margins.

Description: Supp Table 3 (online only). Based on Holotype (♂) MPE00291.

Measurements (mm): BL = 47.0, CW = 6.9, IW = 4.5, ISW = 1.5, B11W = 9.4, B11H = 5.8.

Variation: No significant variation from the holotype was observed.

Ecology: Sigmoria beameri n. sp. was collected from the riparian area around Toby Creek and was found under oak and maple leaves.

Distribution: Only known from the type locality, a small wooded area around Toby Creek, located in the Coastal Plain region of South Carolina.

Etymology: This species is named for its discoverer, Dr. David Beamer of Nash Community College. The specific name is a genitive noun derived as a patronym.