Tomarus pilcopataensis López-García & Deloya, 2019, new species

Zoobank.org/ urn:lsid:zoobank.org:act: AC215467-CC35-4350-8574-00D3BA0652B2 (Figs. 20–22)

Type Material. 16. Holotype labeled “ PERU: / Cuzco; / Pilcopata, / 600 meters / 11-14 xii 1979 / J.B. Heppner / premontane moist forest” (16 USNM).

Description. Holotype. Habitus as in Fig. 20. Length 27.7 mm; width across humeri 14.3 mm. Head: Frons coarsely rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus narrowed towards apex, base width 3.8 times as apex. Clypeal teeth triangular, separated by less than a tooth diameter. Mandibles with 2 apical, acute teeth and a rounded basal lobe. Interocular distance equal to 4.1 times eye width. Antennal club short, subequal in length to antennomeres 2–7. Pronotum: Surface with dense, large punctures. Apical tubercle small, rounded. Fovea shallow, striate, and about 1/5 as wide as interocular distance. Margin of apical angles entire. Elytra: First interval punctate, punctures similar in size to those on other intervals. Pygidium: Surface coarsely rugose. Slightly convex in lateral view. Apex regularly rounded. Legs: Protibia tridentate with an additional inconspicuous, basal convexity. Protarsus subcylindrical, not enlarged; inner claw entire. Metatibia not narrowed before apex, sides nearly parallel; apex slightly crenulate, with 33 spinules. Parameres: Lateral teeth short, widely triangular; apical fourth narrowed, apices directed outwards (Figs. 21–22).

Etymology. The name of the species refers to the type locality, Pilcopata.

Distribution. Tomarus pilcopataensis is known only from Pilcopata, a locality in the premontane moist forest of Peru at an elevation of 600 m (Fig. 23).

Temporal Distribution. December (1).

Diagnosis. Tomarus pilcopataensis is similar to T. pumilus and T. roigjunenti Neita and Ratcliffe, but it can be differentiate by the completely rugose surface of the pygidium (rugose on the basal third in the other two species). The general shape of the parameres of T. pumilus (Fig. 67) is similar that in the new species, but in T. pilcopataensis the apical fourth is strongly narrowed and the lateral medial teeth are wider and not directed backwards (Figs. 21–22).

Comments. There are specimens of Tomarus maternus Prell and Tomarus gyas Erichson with the same collection and locality data (deposited at USNM), which indicates that these three species have sympatric populations in the western Amazonian region. Unfortunately, we were not able to find additional specimens of T. pilcopataensis in three entomological collections in Peru nor during a visit to the type locality in Manu National Park (Cusco).