Nyctiophylax (Paranyctiophylax) nepenthes Oláh & Johanson 2010, new species

Figures 94–96

Diagnosis: This species is similar to N. (P.) icelus Neboiss from Sulawesi, N. (P.) padangensis Malicky from Sumatra, N. (P.) pongdiatensis Malicky & Chantaramongkol from Thailand, N. (P.) simaritensis Malicky & Chantaramongkol from Sumatra, and N. (P.) temburongensis Malicky from Brunei in having slender, ventradcurving ventral paraproctal processes, no sclerotized dorsal paraproctal processes, and a ventroapical median lobe produced posterad from sternite IX. The new species is most similar to N. (P.) icelus, from which it is separated by its long, clavate cerci; gonopods with their apices excised, not pointed; and 6 stout spines on the endotheca.

Description: Male. Body uniformly pale brown. Maxillary palp formula (I,II)-IV-III-V. Forewing brown, with hypertrophied longitudinal vein R and pronounced hyaline patch pattern of 3 hyaline windows: (1) along crossvein r -m, (2) along crossvein m -cu and point of basal forking of M, and (3) on arculus (at junction of A1+A2+A3 and wing margin). Forewing length 3.0 mm; A1, A2, and A3 looping.

Male genitalia. Abdominal segment IX forming robust sternite, nearly ovoid in lateral view (Fig. 94) with well-developed apicoventral mesal lobe; tergum IX membranous, with no narrow sclerotized stripe discernible. Segment X membranous, with pair of long, digitate, hyaline processes weakly discernible in holotype. Cerci setose, narrow, elongate, clavate in lateral view (Fig. 94); almost as long as ventral paraproctal processes. Paraproctal complexes highly sclerotized; each including narrow, paraproctal base and large ventral paraproctal processes curving ventrad and bearing 4–6 setae in row on outer surface at vertical apical half; no sclerotized dorsal paraproctal processes discernible; pair of hyaline digitate processes possibly associated with paraproctal complexes. Gonopods elongate, slender, slightly S-shaped in lateral view (Fig. 94), arching evenly mesad in ventral view (Fig. 95); apices excised in lateral and ventral view; ventrobasal “elbow” absent; base large, bulbous. Phallic apparatus (Fig. 96) located dorsally in genitalia, guided by paraprocts and segment X. Phallotheca thick, oriented horizontally; phallothecal sclerotization ending with distinct edge; apex of phallic apparatus membranous, the retracted membranes covered with granules; retracted membranes with row of 4 anterad-directed, stout spines in phallobase and 2 stout spines distally immediately below horizontally oriented, elongate phallotremal sclerites.

Holotype male: MALAYSIA: Sabah: Tawau, Maliau Basin, Nepenthes Camp, Camel Trophy Hut, 4 ° 43 ’ 59.3 ’’ N, 116 ° 52 ’ 39.7 ’’ E, 999 m, 7.xii.–8.xii.2007, 22W black light [B. Viklund & N. Jönsson]— 1 male (NHMS).

Paratypes: Same data as holotype, except Heli Pad, 4 ° 43 ’ 59.3 ’’ N, 116 ° 52 ’ 42.4 ’’ E, 1003 m, 8.xii.2007, 22W black light [B. Viklund & N. Jönsson]— 1 male (OPC); same data, except crossing stream, 4 ° 43 ’ 58.9 ’’ N, 116 ° 52 ’ 40.7 ’’ E, 994 m, 9.xii.–14.xii.2007, Mega Malaise Trap (6 m long) [B. Viklund & N. Jönsson]— 1 male (NHMS).

Distribution: Malaysia (Sabah).

Etymology: Nepentes, named after the “Nepenthes Camp”, the type locality of the species.