Homoneura (Homoneura) beckeri

Key to all species of the Homoneura (Homoneura) beckeri group in Oriental Region

(Figures for previously described species can be found in the cited references)

1. At least abdominal tergites 1–2 entirely black............................................................... 2

- Abdominal tergites 1–2 mostly yellow..................................................................... 8

2. At least one of abdominal tergites 1–5 with yellow markings................................................... 3

- Abdominal tergites 1–5 entirely deep black................................................................. 5

3. Abdominal tergites 1–4 entirely black (Fig. 38); aedeagus with 3 pair of long sword-like apical processes (Figs. 70–71) H. (H.) ensata sp. nov.

- At least one of abdominal tergites 1–4 with yellow markings; aedeagus not as above................................. 4

4. Upper half of occiput with a large brown spot on each side, extending to top margin of eye; abdominal tergite 3 with narrow yellow anterior margin, tergites 4–5 black on posterior 1/2 and yellow on anterior 1/2; aedeagus with a pair of falciform dorsal sclerites in ventral view (Malloch 1929)............................................... H. (H.) occipitalis Malloch

- Upper half of occiput usually without large brown spot on each side; abdominal tergite 3 yellow anterolaterally; tergites 4–6 each yellow with a blackish brown median spot, which is confluent with blackish brown posterior band in tergites 4–5; aedeagus not as above(Figs. 94–95)........................................................ H. (H.) trilineata sp. nov.

5. Abdominal tergite 6 entirely black................................................ H. (H.) bomiensis Gao & Yang

- Abdominal tergite 6 yellow with brown median spot.......................................................... 6

6. Face yellow with a pair of black spots; surstylus not curved in lateral view; hypandrium without ventral processes (Sasakawa 1992)...................................................................... H. (H.) immaculata (de Meijere)

- Face yellow with a large brown median spot; surstylus curved in lateral view; hypandrium with a pair of broad short ventral processes............................................................................................ 7

7. First flagellomere yellow in basal half and blackish brown in apical half (Fig. 5); mesoscutum with a pair of brownish yellow lateral stripes (Fig. 16)......................................................... H. (H.) dulongjiangica sp. nov

- First flagellomere entirely blackish brown (Fig. 4); mesoscutum without lateral stripes (Fig. 15)...... H. (H.) cornis sp. nov.

8. Face with brown stripe, band or spot......................................................................9

- Face without stripe, band or spot........................................................................ 14

9. Face with one transverse band or two small horizontal spots on ventral 1/3 or near ventral margin..................... 10

- Face with a pair of large brown elliptical spots (sometimes two spots slightly confluent), occupying majority of face...... 11

10. Antennal 1st flagellomere yellow with dark apex; hypandrium slightly concave at middle; aedeagus with a pair of acuate lateral processes in ventral view (Sasakawa 1992)........................................... H. (H.) beckeri (Kertész)

- Antennal 1st flagellomere entirely black; hypandrium projecting at middle; aedeagus without acuate lateral processes in ventral view (Yang, Hu & Zhu 2002).......................................... H. (H.) nigrantennata Yang, Hu & Zhu

11. Abdominal tergite 6 with a pair of blackish brown lateral bands (Fig. 34)................. H. (H.) baihualingensis sp. nov.

- Abdominal tergite 6 without lateral bands................................................................. 12

12. Abdominal tergite 5 with a triangular median spot but lacking a blackish brown transverse band; protandrium with a ventral process; aedeagus consisting of a basal anterior sclerite with a pair of triangular lateral processes, a pair of basally furcated dorsal sclerites convergent at middle and a pair of hammer-shaped dorsal processes in ventral view (Shi & Yang 2009)..................................................................................... H. (H.) convergens Shi & Yang

- Abdominal tergite 5 with a triangular median spot confluent with a blackish brown transverse posterior band; protandrium without ventral process; aedeagus not as above............................................................. 13

13. Mesoscutum with 5 brownish yellow longitudinal stripes (Fig. 14); surstylus consisting of 1 process; aedeagus distinctly incised, with a pair of dorsal sclerites and a pair of preapical lateral processes (Figs. 52–56)......... H. (H.) conica sp. nov.

- Mesoscutum without longitudinal stripes; surstylus consisting of 2 processes; aedeagus not as above (Shi & Yang 2009)................................................................................. H. (H.) convergens Shi & Yang

14. Fore tibia with an incomplete brown ring at basal 1/3.................................. H. (H.) gemmiformis sp. nov.

- Fore tibia without brown ring........................................................................... 15

15. Antennal 1st flagellomere unicolorous.................................................................... 16

- Antennal 1st flagellomere bicolored...................................................................... 19

16. Antennal 1st flagellomere brownish yellow; aedeagus with a pair of acuate lateral processes, a pair of incurved horn–like apical processes in ventral view (Shi & Yang 2009).................................... H. (H.) yinggelingica Shi & Yang

- Antennal 1st flagellomere entirely black; aedeagus not as above................................................ 17

17. Wing immaculate (Fig. 31); protandrium with a long narrow furcated ventral process; surstylus broad hairy digitiform; aedeagus without distinct apical incision, consisting of a pair of dorsal sclerites, with a pair of lateral median teeth, a pair of ventral teeth and a pair of dorsal teeth (Figs. 81–85).......................................... H. (H.) longiprocessa sp. nov.

- Wing with a narrow brown stripe or spot on dm-cu; male genitalia not as above................................... 18

18. Mesoscutum with brown stripe; tergite 6 with a small brown median spot; surstylus with a knife-shaped process in lateral view; aedeagus with a pair of acuate triangular ventral processes and a pair of subuliform dorsal processes in ventral view (Shi & Yang 2009)..................................................................... H. (H.) flavida Shi & Yang

- Mesoscutum without stripe; tergite 6 without spot; surstylus consisting of a broad hairy digitiform process and a narrow curved bar-like process in lateral view; aedeagus with a pair of acuate lateral processes and a pair of wide furcated dorsoapical processes in ventral view (Shi & Yang 2009)........................................... H. (H.) tortifurcata Shi & Yang

19. Wing with a narrow brown stripe or spot on dm-cu.......................................................... 20

- Wing immaculate, without stripe or spot.................................................................. 25

20. Abdominal tergite 6 without spot; hypandrium with a pair of short bar-like ventral processes, aedeagus with a pair of dorsoapical teeth (Shi & Yang 2009)...................................................... H. (H.) apicomata Shi & Yang

- Abdominal tergite 6 with a brown median spot; hypandrium and aedeagus not as above............................. 21

21. Tarsomere 5 entirely yellow to yellowish brown; surstylus undulating (Shi & Yang 2009.... H. (H.) semicircularis Shi & Yang

- Tarsomere 5 partly or entirely brown to black; surstylus not undulating.......................................... 22

22. Tarsomere 5 brownish black; surstylus only with a stick-like process, slightly curved and narrow apically; aedeagus with a pair of curved subuliform processes, a pair of acuate lateral processes and a pair of dorsal teeth on dorsal sclerites (Shi & Yang 2009)........................................................................ H. (H.) nigritarsis Shi & Yang

- Tarsomere 5 dark yellow in basal half and brown in apical half; surstylus consisting of a small subuliform process and a digitifom process; aedeagus with 1-3 pairs of small acuate processes or teeth on dorsal sclerites.......................... 23

23. r-m before middle of discal cell (Fig. 32); abdominal tergite 2 without spot or band (Fig. 41); aedeagus with a pair of slender dorsal sclerites excurved at apex, and a pair of long coniform lateral processes at base (Figs. 89–90).. H. (H.) pilifera sp. nov.

- r-m at middle of discal cell; abdominal tergite 2 with spot or band; aedeagus not as above........................... 24

24. Protandrium with a long T-shaped ventral process; aedeagus with dorsal sclerites curved in lateral view (Shi & Yang 2009)................................................................................. H. (H.) latissima Shi & Yang

- Protandrium with a short V-shaped ventral process; aedeagus with dorsal sclerites straight in lateral view (Li, Li & Yang 2008)......................................................................... H. (H.) yunnanensis Li, Li & Yang

25. Surstylus consisting of 3 processes in lateral view; hypandrium with a pair of long apodemes and a small apodeme at median incision; aedeagus with 3 pairs of preapical dorsal teeth (Figs. 76–80)......................... H. (H.) latizona sp. nov.

- Surstylus consisting of 1 process in lateral view; hypandrium and aedeagus not as above............................ 26

26. Abdominal tergites 3–5 without median spot (Fig. 35); hypandrium without hypandrial apodeme; gonopod incurved at apex in ventral view; aedeagal apodeme short (Figs. 50–51)....................................... H. (H.) brevituba sp. nov.

- Abdominal tergites 3–5 each with a median spot; male genitalia not as above.....................................27

27. Abdominal tergites 3–5 with blackish brown posterior band confluent with triangular median spot; surstylus short claw-like, not projecting beyond tip of epandrium in lateral view; hypandrium without lobate ventral sclerites in ventral view; aedeagus with bifurcated sclerite ventrally and two pairs of spine-like processes on dorsal and ventral membranous parts (Sasakawa 1992)......................................................................... H. (H.) fasciventris Malloch

- Abdominal tergites 3–5 each with a black median spot and a pair of black lateral bands; surstylus long knife-like, projecting beyond tip of epandrium in lateral view; hypandrium with a pair of lobate ventral sclerites in ventral view; aedeagus obliquely truncate at tip with a small acuate process (Shi & Yang 2009).............................. H. (H.) zhangae Shi & Yang