Habetia tuta Ingrisch 2021, sp. nov.

Figs 1B, 3C, 5E–F, 8B, 11A, 12A–C, 18A, 20H, Map 1(3).

Holotype (male): Papua New Guinea: East Sepik, Lager am Rosensee [= Lake Chambri], (4°18’S, 143°7’E), 16.ii.1913, leg. Bürgers (KAE 267)—depository: Museum für Naturkunde Berlin (ZMB).

Other specimens studied: Papua New Guinea: East Sepik, Lager am Töpferfluss [= Keram River], (4°27’S, 144°13’E), 22.iv.1913, leg. Bürgers (KAE 317)— 1 female (ZMB); Quelllager, (4°23’S, 142°47’E), 13.–16.viii.1912, leg. Bürgers (KAE 128)— 1 male (paratype) (ZMB).

Diagnosis. The new species differs from its congeners by the titillators that form a pair of simple curved beams that are never markedly widened and have the apical area almost fully recurved (about 150–160°) against the truncate basal stem and are slightly narrowed but obtuse at tip; while in the other species with simple titillators the apical area is either less strongly recurved and has widened and/or narrowed sections in the apical area (H. pallida sp. nov., H. pedala sp. nov.) or the apical area is recurved against the basal area for little more than 180° and has the tip broadly truncate (H. imitatrix), or the tip of the titillators is dentate (H. dentata sp. nov.). The male cerci have the apico-internal process compressed and at tip divided into a spined dorsal and an obtuse ventral lobe in H. tuta sp. nov. while in H. pallida, that has the most similar cerci, the dorsal lobe is truncate, only half as long as the ventral lobe and not really separated from it, the ventral lobe terminates into an acute tip and the internal surface of the cercus is provided with a little wavy projecting ridge, in H. pedala the apical internal tooth is divided at end into a pair of stout acute teeth of equal length, in H. imitatrix the apical projection of the male cercus is dorso-ventrally compressed and widened and both apical lobes carry a minute spinule at tip, and in H. dentata the apical internal projection of the male cercus is prolonged and strongly modified.

Description. Coloration of face uniformly pale, of general color; lateral areas of face and genae little darker than central area of face, which might be due to conservation; antennal scrobae faintly darkened; mandibles black with few small yellow and dark red spots. Femora with the following number of spines on ventral margins of legs: (1) a 5–8, p 4–8; (2) a 7, p 2–4; (3) a 11–15, p 14–16 (n=3).

Male. Stridulatory file on underside of left tegmen with a distinct step in distal area, in holotype with 66 teeth, of which 7 narrow teeth at base, 36 in main part of file, 5 at a step, 13 clearly separated at end, and 6 of decreasing size at end (Fig. 18A), in paratype with 69 teeth, of which 8 narrow teeth at base, 35 in main part of file, 5 at a step, 13 clearly separated at end, and 6 of decreasing size at end. Mirror behind stridulatory file with straight anterior and posterior margins and convex lateral margins; slightly longer than wide, greatest width anteriorly. Tenth abdominal tergite transverse, in midline furrowed and at fore margin with an obtuse-angular incision filled by membrane, from hind margin with a narrow elongate triangular sub-membranous area; fully sclerotized areas with apical margin obtuse-angular but with rounded angle above bases of cerci (Fig. 8B). Cerci in about basal half roughly cylindrical, in apical half with an obtuse dorsal carina and strongly concave internal surface; at end of cercus, at dorsal margin from nearly tip, at ventral margin from beginning of apical third with a large internal projection with concave proximal surface that is before end divided into a short rounded dorsal process with narrowing margins that ends into an acute black spine, and into a small compressed ventral flap that is curved posteriorly, surpasses the dorsal spine, and carries a minute black spinule at tip (Fig. 12A). Subgenital plate with convex lateral margins that form in basal two thirds a wide, dorso-ventrally compressed rim that frame the strongly convex central area that is dived from base to nearly apical third into a pair of ovoid hemispheres; apical area short and narrow, flattened and with a faint median carina, with two short styli (Fig. 11A); above the subgenital plate there is a semi-tube-shaped sclerotized phallus sheath that holds the phallus when at rest, it has the dorsal margins faintly granular and terminates into a pair of separate rounded lobes (Fig. 8B). Its function is unknown. Titillators simple, laterally compressed, dark or red brown, forming a pair of simple, curved sclerites; at base faintly curved, then straight, followed by an about 150–160° curve, in apical area faintly narrowed, tip rounded (Figs 12B–C). With a pair of spindle-shaped narrow elongate lateral sclerites.

Female. Subgenital plate rhombic with up curved lateral margins that are in basal area strongly prolonged dorsad; at apico-lateral angles with up-bent acute cones (Figs 5E–F). Ovipositor almost twice as long as body; behind basal constriction straight with dorsal valves in basal half faintly higher than in apical half; dorsal margin faintly convex but not raised (Fig. 1B).

Measurements (2 males, 1 female).—Body w wings: male 44–45, female 53; w/o wings: male 33–37, female 39; pronotum: male 9.2–9.5, female 10; tegmen: male 33–34, female 40; hind femur: male 23–24, female 29; ovipositor length: female 69; ovipositor height: female 2.7 mm.

Etymology. The name of the new species refers to the cover encompassing the phallus when at rest; from the Latin adjective tutus, tuta = protected.