16. Tylodinus triumforium Luna-Cozar, sp. nov.

(Figures 23, 26, 56–57, 109, 130)

Diagnosis. Length male, 3.9–4.5 mm, female, 3.5–4.2 mm. Width male, 2.3– 2.1 mm, female 1.9–2.1 mm. Body approximately1.9x longer than wide. Rostrum with small and shallow punctures. Elytra clothed with dark brown to light brown scales on disc, with yellow scales intermixed, denser on sides and in patches on intervals 1–3 at declivity, covering apical part of tubercle on I3 on middle of elytra and I4 on base of declivity, and covering tubercle on I2 almost completely. Male abdomen with ventrite 2 longer than 3–4 combined. Middle and hind tibiae with external tooth. Male genitalia with apical process in ventral view small (Figure 109). Female genitalia with median membranous area of sternite eight as long as basal plate, spermatheca shallowly curved.

Geographic distribution. México (Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1500–2200 m.

Derivation of specific name. Name derived from the Latin word triumphus = triumph, and named after the El Triunfo Biosphere Reserve, the protected natural area where the species was collected.

Material examined. Total 11 males, 13 females. Holotype male (CMNC): México: Chiapas, Mpio Angel Albino Corzo, Reserva Biosfera El Triunfo, Campamento El Quetzal, elev. 2220 m (15.70819°N 92.93070°W), 21.v.2008, coll. R. S. Anderson, mossy oak forest litter. Paratypes: México: Chiapas, Angel Albino Corzo, Reserva Biosfera El Triunfo, Campamento El Quetzal, elev. 2220 m (15.70819°N 92.93070°W), 21.v.2008, coll. R. S. Anderson — 1 female (CMNC); La Concordia, 4 km SE Custepec, elev. 2125 m (15.70658°N, 92.93126°W), 20.v.2008, coll. LLAMA (Wa-A-03-2-29) — 1 male (CMNC); La Concordia, 2 km SE Custepec, elev. 1520 m (15°43'13.58''N, 92°57'4.18''W), 17.v.2008, coll. LLAMA (Wa-A-02-1-39) — 2 males, 1 female (CMNC, ECOS); same locality (15.72051°N, 92.95164°W), LLAMA (Wa-A-02-1-42) — 1 male, 1 female (CMNC, BMNH); same locality, elev. 1700 m (15.71595°N, 92.93842°W), LLAMA (Wa-A-02-2-03) — 1 female (CMNC); same locality (15.71603°N 92.93808°W) LLAMA (Wa-A-02-2-19) — 1 female (ECOS); same locality (15.71580°N 92.93811°W), LLAMA (Wa-A-02-2-24) — 1 male (ECOS); same locality (15.71576°N 92.93812°W), LLAMA (Wa-A-02-2-25) — 1 female (CMNC); same locality, elev. 2125 m (15.70708°N, 92.93131°W), LLAMA (Wa-A- 03-2-17) — 1 male (CMNC); same locality, elev. 2140 m (15.70925°N, 92.92929°W), LLAMA (Wa-A-03-1-48) — 1 female (CWOB); 3.5 km ESE Custepec, Campamento El Quetzal, elev. 1850 m (15°43'.1''N, 92°55'59.9''W), 17.vii.2007, coll. M. G. Branstetter — 1 male, 1 female (CMNC); Reserva de la Biósfera El Triunfo, Campamento El Quetzal, elev. 1830 m (15°43'17''N, 92°56'20.8''W), 20.v.2008, coll. R. S. Anderson — 1 female (CMNC); same locality, 22.v.2008, coll. R. S. Anderson — 1 female (CMNC); same locality, elev. 1530 m, 18.vii.2007, coll. R. S. Anderson — 1 male (CMNC); same locality, elev. 1800 m, 18.vii.2007, coll. R. S. Anderson — 1 female (CMNC); same locality (15.72188°N, 92.93677°W), elev. 1900 m, 19.v.2008, coll. R. S. Anderson — 1 male (CMNC); same locality (15.72178°N, 92.94544°W), elev. 1700 m, 17.v.2008, coll. R. S. Anderson — 2 females (CMNC, ECOS); same locality, elev. 2177 m, 23.v.2008, coll. J. L. Cozar — 1 male (CMNC).