1 Mainland species; aedeagus symmetrical (Fig. 65) ………………………………………………...……. 2

1’ Island species; aedeagus asymmetrical (Fig. 58) ……………………………...………………………. 5

2 General color dark brown ………………………. 3

2’ General color yellow or pale brown.………….. 4

3 Tergum 10 with U-shaped sclerite; vesicle broadly arcuate, shorter than wide; aedeagus with dorsolateral pair of narrow sclerites; known from Vietnam.………………………………….. malickyi

3’ Tergum 10 apparently without U-shaped sclerite; vesicle longer than wide; known from Yunnan, China ……………………………………... nigrifulva

4 Cercal segments with enlarged dorsal bristles on mid segments (Fig. 66); vesicle broadly arcuate, shorter than wide (Fig. 64); ventroapical aedeagal lobes sparsely armed laterally with fine setal-like spines, apex bilobed (Fig. 65) …………….. spinosa

4’ Cercal segments without conspicuously enlarged dorsal bristles; vesicle about as wide as long (Fig. 69); ventroapical aedeagal lobes armed laterally with dense patch of fine setal-like spines, apex indistinctly lobed (Fig. 70) ……………….. swanni

5 Known from the Philippines …………………….. 6

5’ Known from Borneo …………………………….. 7

6 Sinistral aedeagal lobe weakly sclerotized along ventrolateral margin (Fig. 52); aedeagal lobes armed with small spines and microtrichia (Fig. 55) ………………………………………………. cebuano

6’ Sinistral aedeagal lobe membranous; aedeagal lobes armed with apical patch of cultriform spines ………………………………………….. mindanensis

7 Sinistral aedeagal lobe bifurcate apically and armed with a few large spines at dorsolateral margin (Fig. 46) …………………………... anomala

7’ Sinistral aedeagal lobe trifurcate apically and armed dorsolaterally with small spicules; known from Mt. Kinabalu ………………………… concolor