Swammerdamella marginata Cook

(Figs. 5, 6, 21, 22, and 57–61)

Swammerdamella marginata Cook, 1956b: 24, figs. 1D (male terminalia), 2D (male segment 7), 3G (wing), 4E (male segment 6), 5H

(female terminalia). Type locality: United States, New Mexico, Las Vegas. Distr.– United States (New Mexico), Mexico (Guadalajara).

Ref.– Cook, 1967: 4 (Nearctic catalog). Holotype ♂, NMNH.

Diagnosis. Male tergite 6 large, produced posteriorly into a broad, curved lobe, sternite 6 ornamented, more shallowly incised posteriorly.

Material Examined. 4 ♂♂, 2 ♀♀, United States, California, Los Angeles Co., Los Angeles, Exposition Park, 34.018°N 118.288°W, BioSCAN site 1, 28 March–4 April 2014, Malaise trap, 54 m, Coll. Lisa Gonzalez and Dean Pentcheff, BioSCAN sample 15775; 1 ♂, same data, but 1–8 April 2016, BioSCAN sample 18198; 1 ♀, same data, but BioSCAN sample 16170; 3 ♀♀, same data, but BioSCAN sample 16029; 1 ♀, same data, but 3–10 June 2016, BioSCAN sample 18273; 3 ♂♂, 8 ♀♀, same data, but 2–9 May 2014, BioSCAN sample 15915 (1♀ on slide); 1 ♀, same data, but BioSCAN sample 18373; 8 ♂♂, 1 ♀, United States, California, Los Angeles Co., Los Angeles, University Park, 34.034°N 118.281°W, BioSCAN site 3, 2–9 April 2014, Malaise trap, 63 m, Coll. Peggy Hentschke, BioSCAN sample 15897; 1 ♀, same data, but 3–10 December 2014, BioSCAN sample 16969; 1 ♂, 1 ♀, same data, but BioSCAN sample 16251; 1 ♀, same data, but BioSCAN sample 16072; 1 ♀, same data, but 26 February–5 March 2014, BioSCAN sample 15721; 2 ♂♂, 2 ♀♀, same data, but 30 April–7 May 2014, BioSCAN sample 16068; 1 ♀, same data, but 29 October–5 November 2014, BioSCAN sample 16774; 1 ♀, same data, but BioSCAN sample 16603; 1 ♂, United States, California, Los Angeles Co., Los Angeles, Los Feliz, 34.118°N 118.284°W, BioSCAN site 4, 29 March–4 April 2014, Malaise trap, 220 m, Coll. Mark and Jane Pisano, BioSCAN sample 15950; 1 ♂, 3 ♀♀, same data, but 1–15 March 2014, BioSCAN sample 15757; 1 ♀, same data, but BioSCAN sample 16808; 4 ♀♀, same data, but BioSCAN sample 16632; 10 ♂♂, 40 ♀♀, United States, California, Los Angeles Co., Los Angeles, Silverlake, 34.093°N 118.274°W, BioSCAN site 5, 29 March–5 April 2014, Malaise trap, 166.1 m, Coll. Walter Renwick, BioSCAN sample 15916; 2 ♂♂, 2 ♀♀, same data, but 1–8 February 2014, BioSCAN sample 15589; 1 ♂, 31 ♀, same data, but BioSCAN sample 16268; 2 ♂♂, 42 ♀♀, same data, but BioSCAN sample 16113; 2 ♂♂, 33 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15735; 2 ♂♂, 3 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16109; 1 ♂, 66 ♀♀, same data, but 2–9 November 2013, BioSCAN sample 15318; 5 ♀♀, same data, but 28 September–5 October 2013, BioSCAN sample 15226; 4 ♂♂, United States, California, Los Angeles Co., Los Angeles, Los Feliz, 34.116°N 118.279°W, BioSCAN site 6, 1–8 February 2014, Malaise trap, 199 m, Coll. Jeff, Aaron and Jacob Koch, BioSCAN sample 15572; 3 ♀♀, same data, but BioSCAN sample 16107; 1 ♂, same data, but 1–9 March 2014, BioSCAN sample 15746; 2 ♀♀, same data, but 4–10 May 2014, BioSCAN sample 16104; 1 ♂, same data, but 15– 20 September 2013, BioSCAN sample 15207; 2 ♀♀, same data, but 29 March–14 April 2014, BioSCAN sample 15939; 1 ♂, United States, California, Los Angeles Co., Los Angeles, Silverlake, 34.102°N 118.257°W, BioSCAN site 7, 29 March–5 April 2014, Malaise trap, 158.9 m, Coll. Joe Hogg, BioSCAN sample 15834; 1 ♀, same data, but BioSCAN sample 16193; 1 ♀, same data, but BioSCAN sample 16189; 2 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15830; 1 ♀, same data, but 26 April–10 May 2014, BioSCAN sample 16003; 2 ♂♂, United States, California, Los Angeles Co., Los Angeles, Elysian Valley, 34.103°N 118.243°W, BioSCAN site 9, 29 March–5 April 2014, Malaise trap, 106.7 m, Coll. John Rodriguez, BioSCAN sample 15877; 1 ♂, same data, but 28 December 2013 – 4 January 2014, BioSCAN sample 15498; 10 ♂♂, 25 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15700; 2 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16050; 1 ♀, same data, but 28 September–5 October 2013, BioSCAN sample 15274; 1 ♀, same data, but 2–9 November 2013, BioSCAN sample 15279; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Leimert Park, 34.014°N 118.321°W, BioSCAN site 10, 28 September–5 October 2013, Malaise trap, 38 m, Coll. LaChristian Steptoe, BioSCAN sample 15220; 1 ♂, 13 ♀♀, United States, California, Los Angeles Co., Los Angeles, Korea Town, 34.072°N 118.291°W, BioSCAN site 11, 5–12 April 2014, Malaise trap, 85.6 m, Coll. Peter Ralph, BioSCAN sample 16056; 2 ♀♀, same data, but 2–9 March 2014, BioSCAN sample 15796; 2 ♂♂, 4 ♀♀, same data, but 4–13 May 2014, BioSCAN sample 16060; 1 ♀, same data, but BioSCAN sample 16482; 5 ♂♂, United States, California, Los Angeles Co., Los Angeles, Mid-City, 34.047°N 118.334°W, BioSCAN site 12, 29 January–5 February 2014, Malaise trap, 52.6 m, Coll. Sharon Oxborough, BioSCAN sample 15560; 4 ♂♂, 1 ♀, same data, but 1–8 February 2014, BioSCAN sample 15600; 4 ♀♀, same data, but BioSCAN sample 16323; 1 ♀, same data, but BioSCAN sample 16142; 4 ♂♂, 13 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15784; 3 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 15948; 3 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16838; 2 ♀♀, same data, but 14–21 September 2013, BioSCAN sample 15202 (on slide); 2 ♂♂, 16 ♀♀, same data, but 29 March–5 April 2014, BioSCAN sample 15943; 3 ♂♂, United States, California, Los Angeles Co., Glendale, 34.149°N 118.218°W, BioSCAN site 13, 28 March–5 April 2014, Malaise trap, 245.4 m, Coll. Karen Hoffman, BioSCAN sample 15851; 1 ♂, 1 ♀, same data, but BioSCAN sample 16381; 1 ♂, 1 ♀, same data, but BioSCAN sample 16211; 1 ♂, 4 ♀♀, same data, but BioSCAN sample 16207; 3 ♂♂, 1 ♀, same data, but 1–8 March 2014, BioSCAN sample 15680; 1 ♂, same data, but 3 May 2014 – 9 May 2014, BioSCAN sample 15998; 1 ♂, same data, but 29 November–6 December 2014, BioSCAN sample 16893; 3 ♂♂, United States, California, Los Angeles Co., Los Angeles, Gardena, 33.876°N 118.288°W, BioSCAN site 15, 29 March–5 April 2014, Malaise trap, 9 m, Coll. Betty Defibaugh, BioSCAN sample 15903; 12 ♀♀, United States, California, Los Angeles Co., Los Angeles, Hollywood, 34.095°N 118.334°W, BioSCAN site 16, 2–9 April 2014, Malaise trap, 100 m, Coll. Tony Hein, BioSCAN sample 15980; 1 ♀, same data, but BioSCAN sample 16345; 1 ♂, same data, but 29 January–5 February 2014, BioSCAN sample 15611; 5 ♀♀, same data, but 1–18 January 2014, BioSCAN sample 15608; 7 ♀♀, same data, but BioSCAN sample 16341; 3 ♀♀, same data, but BioSCAN sample 16137; 13 ♀♀, same data, but 26 February–5 March 2014, BioSCAN sample 15801; 4 ♀♀, same data, but 30 April–7 May 2014, BioSCAN sample 15984; 12 ♀♀, same data, but 29 October–5 November 2014, BioSCAN sample 16690; 1 ♀, same data, but BioSCAN sample 16686; 1 ♂, 1 ♀, same data, but BioSCAN sample 16519; 23 ♂♂, 9 ♀♀, United States, California, Los Angeles Co., Glendale, 34.159°N 118.247°W, BioSCAN site 17, 29 March–5 April 2014, Malaise trap, 190 m, Coll. Celeste Armstrong, BioSCAN sample 15911; 5 ♀♀, United States, California, Los Angeles Co., Glendale, 34.159°N 118.247°W, BioSCAN site 17, 1–8 February 2014, Malaise trap, 190 m, Coll. Celeste Armstrong, BioSCAN sample 15670; 1 ♀, same data, but BioSCAN sample 16200; 2 ♀♀, same data, but BioSCAN sample 16196; 6 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15674; 5 ♂♂, 4 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16013; 2 ♂♂, 10 ♀♀, United States, California, Los Angeles Co., Los Angeles, Larchmont, 34.077°N 118.32°W, BioSCAN site 18, 29 March–5 April 2014, Malaise trap, 76 m, Coll. K.T.Wiegman, BioSCAN sample 15968; 3 ♀♀, same data, but 2 December 2014 – 6 December 2014, BioSCAN sample 16847; 1 ♂, 10 ♀♀, same data, but 1 February 2014 – 10 February 2014, BioSCAN sample 15617; 4 ♀♀, same data, but BioSCAN sample 16305; 1 ♂, 1 ♀, same data, but BioSCAN sample 16127; 3 ♂♂, 45 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15778; 1 ♂, 3 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 15973; 1 ♂, 2 ♀♀, same data, but 2–9 November 2013, BioSCAN sample 15322; 4 ♂♂, 7 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16843; 3 ♂♂, same data, but BioSCAN sample 16501; 26 ♂♂, 96 ♀♀, United States, California, Los Angeles Co., Los Angeles, Carthay, 34.059°N 118.369°W, BioSCAN site 19, 1–8 April 2014, Malaise trap, 41.1 m, Coll. Teresa Dahl, BioSCAN sample 15812 (2♀♀ on slide); 6 ♀♀, same data, but BioSCAN sample 16338 (3 ♀♀ on slide); 2 ♂♂, 11 ♀♀, same data, but 2–9 December 2014, BioSCAN sample 16876; 1 ♂, 3 ♀♀, same data, but 28 January–4 February 2014, BioSCAN sample 15593; 1 ♂, 3 ♀♀, same data, but BioSCAN sample 16334; 2 ♂♂, 10 ♀♀, same data, but BioSCAN sample 16147; 4 ♂♂, 225 ♀♀, same data, but 25 February–4 March 2014, BioSCAN sample 15807; 3 ♂♂, 17 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 15977; 1 ♂, 7 ♀♀, same data, but 28 October–4 November 2014, BioSCAN sample 16871; 1 ♂, 4 ♀♀, same data, but BioSCAN sample 16868; 1 ♀, same data, but BioSCAN sample 16513; 1 ♀, United States, California, Los Angeles Co., Burbank, 34.17°N 118.308°W, BioSCAN site 21, 1–8 March 2014, Malaise trap, 158 m, Coll. Chris Cianci, BioSCAN sample 15668; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Echo Park, 34.074°N 118.264°W, BioSCAN site 22, Malaise trap, 133.5 m, Coll. Erin Johnson, BioSCAN sample 16302; 1 ♀, same data, but 28 January–4 February 2014, BioSCAN sample 15577; 1 ♀, same data, but BioSCAN sample 16298; 2 ♀♀, same data, but 25 February–4 March 2014, BioSCAN sample 15770; 10 ♀♀, United States, California, Los Angeles Co., Los Angeles, Atwater Village, 34.114°N 118.251°W, BioSCAN site 23, 2–10 April 2014, Malaise trap, 119 m, Coll. Sidney Higgins, BioSCAN sample 15858 (1 ♀ on slide); 1 ♀, same data, but 2–8 January 2014, BioSCAN sample 15540; 1 ♀, same data, but BioSCAN sample 16355; 5 ♀♀, same data, but 26 February–5 March 2014, BioSCAN sample 15663; 1 ♀, same data, but 30 April–7 May 2014, BioSCAN sample 16172; 1 ♀, same data, but 29 October–7 November 2014, BioSCAN sample 16707; 3 ♀♀, United States, California, Los Angeles Co., Los Angeles, Eagle Rock, 34.136°N 118.194°W, BioSCAN site 24, Malaise trap, 215 m, Coll. Jim Hogue, BioSCAN sample 16187; 1 ♂, same data, but 1–8 March 2014, BioSCAN sample 15842; 2 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16018; 1 ♂, United States, California, Los Angeles Co., Los Angeles, Pico–Union, 34.046°N 118.275°W, BioSCAN site 25, 29 March–5 April 2014, Malaise trap, 70 m, Coll. Humberto Marquez, BioSCAN sample 15882; 3 ♀♀, United States, California, Los Angeles Co., Los Angeles, Mid–Wilshire, 34.058°N 118.328°W, BioSCAN site 26, 1–8 February 2014, Malaise trap, 58.7 m, Coll. Patty Lombard, BioSCAN sample 15606; 1 ♂, same data, but BioSCAN sample 16663; 3 ♂♂, 13 ♀♀, United States, California, Los Angeles Co., Los Angeles, Eagle Rock, 34.129°N 118.215°W, BioSCAN site 28, 29 March–5 April 2014, Malaise trap, 160.7 m, Coll. Eric Keller, BioSCAN sample 15871; 6 ♀♀, same data, but BioSCAN sample 16418; 3 ♀♀, same data, but 1–8 February 2014, BioSCAN sample 15710; 1 ♀, same data, but 28 December 2013 – 4 January 2014, BioSCAN sample 15480; 1 ♂, 4 ♀♀, same data, but BioSCAN sample 16245; 2 ♂♂, 5 ♀♀, same data, but BioSCAN sample 16241; 3 ♂♂, 6 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15714; 14 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16044; 4 ♂♂, 22 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16755; 6 ♀♀, same data, but 29 November–6 December 2014, BioSCAN sample 16962; 3 ♂♂, 17 ♀♀, same data, but BioSCAN sample 16601; 2 ♂♂, 18 ♀♀, same data, but BioSCAN sample 16597; 1 ♂, 1 ♀, United States, California, Los Angeles Co., Los Angeles, Highland Park, 34.123°N 118.193°W, BioSCAN site 29, 29 March–5 April 2014, Malaise trap, 205.6 m, Coll. Natalie Brejcha, BioSCAN sample 15865; 2 ♀♀, same data, but 21–28 December 2013, BioSCAN sample 15488; 1 ♂, same data, but 1–8 February 2014, BioSCAN sample 15703; 1 ♀, same data, but BioSCAN sample 16233; 3 ♀♀, same data, but BioSCAN sample 16229; 6 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15707; 2 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16032 (1♀ on slide); 1 ♀, same data, but 25 October–8 November 2014, BioSCAN sample 16786; 6 ♂♂, 8 ♀♀, United States, California, Los Angeles Co., Los Angeles, Highland Park, 34.125°N 118.189°W, BioSCAN site 30, 31 March–8 April 2014, Malaise trap, 227.8 m, Coll. Candace Franco, BioSCAN sample 15887; 1 ♂, 4 ♀♀, same data, but BioSCAN sample 16423; 25 ♂♂, 16 ♀♀, same data, but BioSCAN sample 16239; 12 ♂♂, 30, same data, but BioSCAN sample 16235; 10 ♂♂, 4 ♀♀, same data, but BioSCAN sample 15767; 49 ♂♂, 23 ♀♀, same data, but 3–12 May 2014, BioSCAN sample 16038; 13 ♂♂, 2 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16762; 1 ♀, same data, but 29 November–6 December 2014, BioSCAN sample 16955; 5 ♂♂, same data, but BioSCAN sample 16595; 1 ♀, United States, California, Los Angeles Co., Los Angeles, 34.103°N – 118.216°W, BioSCAN site 31, Malaise trap, 74 m, Coll. Julian Donahue, BioSCAN sample 16206; 1 ♀, same data, but BioSCAN sample 16202; 1 ♀, same data, but 1–8 November 2014, BioSCAN sample 16721; 3 ♀♀, United States, California, Los Angeles Co., Los Angeles, Pasadena, 34.163°N 118.118°W, BioSCAN site 33, 1–8 August 2016, Malaise trap, 280 m, Coll. Leslie Harris, BioSCAN sample 18235; 1 ♂, same data, but 1–8 February 2016, BioSCAN sample 18180; 1 ♀, same data, but 1–8 June 2016, BioSCAN sample 18325; 15 ♀♀, same data, but BioSCAN sample 18236; 2 ♂♂, United States, California, Los Angeles Co., Los Angeles, Rosemead, 34.05°N 118.108°W, BioSCAN site 34, 1–8 April 2016, Malaise trap, 135 m, Coll. Deniece Lascano, BioSCAN sample 18230; 2 ♀♀, same data, but 1–8 July 2016, BioSCAN sample 18328; 1 ♀, same data, but 8–25 March 2016, BioSCAN sample 18185; 1 ♀, United States, California, Los Angeles Co., Monrovia. 34.158°N 117.999°W, BioSCAN site 35, 1–8 April 2016, Malaise trap, 214 m, Coll. Brian Brown, BioSCAN sample 18197; 12 ♀♀, same data, but 4–11 August 2016, BioSCAN sample 18370; 3 ♂♂, 7 ♀♀, same data, but 1–15 June 2016, BioSCAN sample 18277; 4 ♀♀, same data, but 1–8 March 2016, BioSCAN sample 18155; 3 ♂♂, 1 ♀, same data, but 1–8 May 2016, BioSCAN sample 18212; 8 ♀♀, same data, but 1–8 October 2016, BioSCAN sample 18422; 1 ♂, 1 ♀, same data, but 1–8 September 2016, BioSCAN sample 18375; 1 ♂, United States, California, Los Angeles Co., San Dimas, 34.086°N 117.822°W, BioSCAN site 36, 1–8 August 2016, Malaise trap, 298 m, Coll. Carla Zorko, BioSCAN sample 18401; 2 ♀♀, same data, but 1–8 June 2016, BioSCAN sample 18341; 3 ♀♀, same data, but BioSCAN sample 18246; 3 ♂♂, 16 ♀♀, United States, California, Los Angeles Co., Claremont, 34.09°N 117.711°W, BioSCAN site 37, 1–8 April 2016, Malaise trap, 341 m, Coll. Kathryn Turner, BioSCAN sample 18249; 1 ♀, same data, but 1–8 July 2016, BioSCAN sample 18338; 1 ♂, 3 ♀♀, same data, but 1–8 June 2016, BioSCAN sample 18337; 7 ♂♂, same data, but 1–8 March 2016, BioSCAN sample 18164; 2 ♀♀, United States, California, San Bernardino Co., Rancho Cucamonga, 34.139°N 117.521°W, BioSCAN site 38, 1–8 August 2016, Malaise trap, 451 m, Coll. Lucila Tagliani, BioSCAN sample 18407; 1 ♂, United States, California, San Bernardino Co., Grand Terrace, 34.024°N 117.315°W, BioSCAN site 39, 7–14 April 2016, Malaise trap, 322 m, Coll. Dave Goodward, BioSCAN sample 18221; 1 ♀, United States, California, Los Angeles Co., El Segundo. 33.925°N 118.425°W, BioSCAN site 40, Malaise trap, 35 m, Coll. Jack Fulmer, BioSCAN sample 18420; 3 ♀♀, United States, California, Los Angeles Co., La Mirada, 33.914°N 118.005°W, BioSCAN site 43, 1–8 August 2016, Malaise trap, 63 m, Coll. Carmen Gendusa, BioSCAN sample 18383; 1 ♀, same data, but 1–8 February 2016, BioSCAN sample 18146; 1 ♂, 2 ♀♀, same data, but 1–8 March 2016, BioSCAN sample 18147; 1 ♂, 1 ♀, same data, but 1–8 June 2016, BioSCAN sample 18321; 1 ♂, 3 ♀♀, same data, but 1–8 April 2016, BioSCAN sample 18265; 1 ♂, 3 ♀♀, same data, but 1–8 October 2016, BioSCAN sample 18385; 1 ♂, 1 ♀, United States, California, Los Angeles Co., Walnut. 33.998°N 117.842°W, BioSCAN site 44, 1–8 April 2016, Malaise trap, 189 m, Coll. Emilia Park, BioSCAN sample 18251; 1 ♀, same data, but BioSCAN sample 18416; 5 ♂♂, 5 ♀♀, United States, California, San Bernardino Co., Chino Hills, 34.003°N 117.734°W, BioSCAN site 45, 1–8 April 2016, Malaise trap, 225 m, Coll. Joe Williams, BioSCAN sample 18253; 1 ♀, same data, but 1–8 June 2016, BioSCAN sample 18339; 8 ♂♂, 3 ♀♀, same data, but BioSCAN sample 18254; 1 ♀, United States, California, Riverside Co., Norco. 33.912°N 117.57°W, BioSCAN site 46, 1–8 April 2016, Malaise trap, 180 m, Coll. Sara Taylor, BioSCAN sample 18227; 1 ♀, same data, but 1–8 August 2016, BioSCAN sample 18392; 12 ♀♀, same data, but 1–8 July 2016, BioSCAN sample 18336; 12 ♀♀, same data, but BioSCAN sample 18335; 1 ♀, same data, but BioSCAN sample 18228; 1 ♀, same data, but BioSCAN sample 18393; 2 ♂♂, 1 ♀, United States, California, Riverside Co., Riverside, 33.984°N 117.313°W, BioSCAN site 47, 1–8 April 2016, Malaise trap, 375 m, Coll. Justin Betz, BioSCAN sample 18223; 2 ♀♀, same data, but 1–8 June 2016, BioSCAN sample 18279 (1 ♀, BioSCAN sample 16032, 2♂♂, 2♀♀, BioSCAN sample 15589 at MZUSP, all other specimens at LACM).

Distribution. United States (southern California and Nevada), Mexico (Guadalajara).

Remarks. Swammerdamella marginata is a very small scatopsid, ranging from 1.30 to 1.40 mm. The type locality of S. marginata is Las Vegas, New Mexico. Cook (1965a) also referred to specimens of this species from additional sites in southern California (San Bernardino, Davis, Riverside, Los Angeles etc.) and Nevada (Reno). The record for Guadalajara, Mexico, may be questionable, but we did not examine the original specimens identified by Cook (1965a).

This was the most abundant species in our samples, with up to 66.1% of the total number of scatopsid specimens of the project: a total of 367 males and 1,315 females were identified in the sorted samples. The species appeared at 39 of the 47 BioSCAN collecting sites, in a total of 182 collecting events. The most abundant collecting sites for S. marginata were 5, 18, 19, 28, and 30. As happened with Quateiella quatei, there is a clear peak of abundance of S. marginata in early-mid-spring, with a second peak in mid-autumn.

There is no information available on the biology of S. marginata. In North America, there are records of adults of S. pygmaea (Loew) associated to flowers of Staphylea floralis (Staphyleaceae) (Cook 1956b). Staphylea has species native to eastern North America and to California (USDA 2019), but no connection so far of this plant genus with S. marginata has been formally proposed. In South America, there is a record of emergence of a species of a Swammerdamella over piles of rotting grass (D.S.A., personal observation). The BOLDSystems has one single specimen of Swammerdamella sequenced for Australia, most certainly not S. marginata.