† Lepiceratus ankylosaurus Jałoszynski, Luo, Yamamoto & Beutel, sp. nov.

(Figs 1–3)

Diagnosis. As for genus.

Description. Body (Figs 1A–D, 2A, B, D, 3A, B, E) stout, length 1.71 mm, region from anterior prothorax to posterior third of elytra approximately parallel-sided; pigmentation light brown with diffuse pattern of darker areas on most convex structures (pronotal and elytral tubercles and ridges); body surface coarsely sculptured and matte. Entire body covered with thin crust of presumably mud particles and glandular secretion, best visible in concave areas on pronotum and elytra and on ventral body side.

Head (Figs 1A–D; 2A–D; 3A–C, E) strongly transverse, length 0.21 mm, width 0.63 mm; with very large and strongly convex compound eyes, in dorsal view occupying nearly entire lateral margin of head; tempora vestigial; vertex and frons convex medially and impressed sub-laterally, with paired oblique dorsal protuberances or ridges.

Antennae short, length 0.45 mm; scapus (Figs 2C, 3C) large, elongate; pedicellus (Figs 2C, 3C) sub-cylindrical, about as broad as long; antennomere 3 (Figs 2C, 3C) about as long as broad; antennomeres 4–6 (Figs 2C, 3C) each slightly broader than 3 and slightly transverse; antennal club (Figs 2C, 3C) slightly flattened dorsoventrally, slightly shorter than all proximal antennomeres combined, with anterior margin nearly straight and posterior margin slightly rounded; oblique apical surface with densely arranged short setae.

Pronotum (Fig. 2A, C) about 0.7 as broad as long, length 0.55 mm, width 0.79 mm; anterior margin sub- trapezoidal and forming short blunt projection overhanging posterodorsal region of head; anterior margin between anterior projection and anterior corner slightly concave; lateral margins irregularly undulate and convergent posterad; posterior margin shallowly and broadly emarginate. Antennal pocket (Figs 1D, 3C) behind each eye open anteroventrally, deep, large and sharply demarcated, with its concave surface smooth. Pronotal disc with convex median area bearing longitudinal rows of several tubercles of various diameters and heights, elongate median region impressed and relatively smooth; impressed lateral portions of pronotal disc with sub-median transverse row of diffuse and weakly elevated tubercles. Setae on pronotum indiscernible.

Elytra (Figs 1A–D, 2A, B, D, 3A, B, E) broadest shortly behind humeri; length 0.95 mm, width 0.91 mm. In lateral view (Figs 2B, 3B) dorsal surface of elytra nearly parallel to longitudinal body axis in anterior half, in posterior half strongly rounded, thus elytral apex not visible in dorsal view. Humeral angles distinctly projecting anterad, anterior margin of each elytron thus slightly concave. Elytral tubercles of various heights and diameters, longest tubercles situated on posterior elytral surface and directed posterad; most tubercles bear one apical bristle (best visible on posterior half of elytra). In lateral view, sub-median ‘tooth’ of elytral edge obtuseangled, concavity behind it rounded.

Three pairs of legs similar. Procoxae (Fig. 1D) triangular, anterior portion separated by prosternal process, posteromesal edges nearly adjacent at midline. Articulation of procoxae with prosternum situated laterally, with notch in nearly straight anterior coxal margin and small sternal condyle. Mesocoxae (Fig. 1D) widely separated and obliquely inserted, flat and distinctly smaller than procoxae; anterior and posterior margins rounded, posterior edge slightly overlapping with trochanteral base. Metacoxae (Fig. 1D) flat, more transverse but otherwise similar to mesocoxae, separated by posteromedian katepisternal excavation, which is narrower than each metacoxa. Protrochanters (Fig. 1D) triangular and about half as long as procoxae. Mesotrochanters (Fig. 1D) slightly smaller. Metatrochanters (Fig. 1D) about as large as protrochanters. Profemora (Fig. 1D) about twice as long as procoxae; their anterior margins slightly curved and ventral margins slightly concave. Meso- and metafemora (Fig. 1D) slightly shorter than profemora, fitting into elytral excavations (Figs 1D, 2D). Protibia (Fig. 1D) sub-cylindrical, shorter than profemora, with fine spines along anterior and posterior margins, lacking distal tibial spur but with posterolaterally directed robust spine at apex. Meso- and metatibiae (Fig. 1D) similar to protibiae. Protarsus (Fig. 1D) including claw slightly shorter than protibia; showing indistinct subdivision into four incompletely delimited subunits and with spines along anterior and posterior edges; single curved claw about as long as apical protarsal subunit. Meso- and metatarsi (Fig. 1D) gradually tapering distad, not showing discernible subdivision.

Abdomen (Figs 1B, D, 2D, 3E) covered with punctures, setae indiscernible.

Derivation of name. After ankylosaurs, the dinosaurs with longitudinal rows of armour plates resembling the rows of tubercles in the new species (noun in apposition).

Stratigraphic range. Late Albian–early Cenomanian.

Hukawng Valley.