Liphistius nawngau Zhan & Xu sp. nov.

Figs 7, 8

Type material.

Holotype: Myanmar ♂, Shan State, Kyaukme Dist., Nawnghkio Township, Nawng Au Vill.; 22.26 ° N, 96.83 ° E, alt. 1096 m; 26. 07. 2019; D. Li et al. leg.; XUX-2019-054A. Paratypes: 9 ♂♂ 3 ♀♀, same data as for holotype; XUX-2019-054 / 054 A / 055 / 055 A / 055 B / 055 D / 056 / 056 A / 057 / 058 / 059 / 060; 5 ♂♂ 2 ♀♀, same township as for holotype; 22.30 ° N, 96.73 ° E, alt. 845 m; 26. 07. 2019 and 14. 07. 2018; XUX-2019-049 / 051 / 052 / 053, XUX-2018-116 / 119 / 123.

Diagnosis.

The male of L. nawngau sp. nov. resembles those of L. lordae and L. pyinoolwin in having an adpressed proximal tegular margin (Fig. 7 C, F), but it can be distinguished from them by the tegulum (T), which bears a distinct transversal ridge in retrolateral view (Fig. 7 C, F vs lacking transversal ridge, figs 5, 6 in Schwendinger 1990; fig. 4 F in Xu et al. 2021), and by the embolus (E), which has smooth longitudinal ridges reaching tip (Fig. 7 E vs having several denticles along longitudinal ridges, fig. 4 F in Xu et al. 2021). The female of L. nawngau sp. nov. can be distinguished from that of L. hpruso by having the posterior stalk triangular (PS) (Fig. 8 vs nearly oval, fig. 3 B – E in Aung et al. 2019).

Description.

Male holotype. Carapace brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 2–6 th larger than others, 5 th largest; chelicerae robust, promargin of cheliceral groove with 11 denticles of variable size; labium and sternum yellowish brown, sternum with few short setae on anterior tip and many long setae on elongated posterior tip; legs yellowish brown, with strong setae and spines. Measurements: BL 15.47, CL 6.63, CW 6.57, OL 7.73, OW 5.56; eye sizes and interdistances: AME 0.08, ALE 0.72, PME 0.35, PLE 0.49, AME – AME 0.08, AME – ALE 0.20, PME – PME 0.08, PME – PLE 0.07, ALE – PLE 0.81, ALE – ALE 0.11, PLE – PLE 0.16, AME – PME 0.12; labium 0.81 long and 0.58 wide; sternum 2.59 long and 1.01 wide; legs: I 19.58 (5.67, 2.78, 4.03, 4.68, 2.42), II 20.42 (5.66, 2.78, 4.13, 4.96, 2.89), III 22.62 (5.96, 2.63, 4.54, 6.29, 3.20), IV 28.26 (7.16, 2.75, 5.82, 8.55, 3.98).

Palp: tibial apophysis (TiA) with four setae of similar length (Fig. 7 A – C); paracymbium (PC) with several setae situated on tip (Fig. 7 A – C); cumulus (Cu) slightly elevated, with 8 tapering spines (Fig. 7 A – C); subtegulum (ST) with weakly developed apophysis (Fig. 7 B, F); contrategulum (CT) with an arched projection distally, and triangular process with wide base (Fig. 7 D); tegulum (T) with distinct transversal ridge in retrolateral view (Fig. 7 C, F); paraembolic plate (PeP) narrow, sclerotized (Fig. 7 A, F); embolus (E) with several longitudinal ridges reaching tip (Fig. 7 E, F).

Female paratype (XUX- 2019-053). Carapace yellowish brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, with gray patches, 2–6 th larger than others, 5 th largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 12 denticles of variable size; labium and sternum yellowish brown; legs with strong setae and spines. Measurements: BL 14.24, CL 6.10, CW 5.30, OL 7.12, OW 4.96; eye sizes and interdistances: AME 0.08, ALE 0.57, PME 0.31, PLE 0.45, AME – AME 0.10, AME – ALE 0.17, PME – PME 0.09, PME – PLE 0.08, ALE – PLE 0.09, ALE – ALE 0.10, PLE – PLE 0.46, AME – PME 0.06; labium 1.23 long and 0.77 wide; sternum 2.68 long and 1.13 wide; palp 10.42 (3.63, 1.84, 2.77, 2.18), leg I 13.33 (4.16, 2.15, 2.90, 2.58, 1.54), II 16.85 (4.35, 2.26, 2.84, 2.84, 1.72), III 15.04 (4.37, 2.35, 2.90, 3.31, 2.11), IV 20.39 (5.92, 1.92, 4.48, 5.33, 2.74).

Vulva: poreplate (PPl) with pair of large anterior lobes and pair of relatively small anterolateral lobes; central dorsal opening (CDO) located at center of poreplate (PPl); receptacular cluster (RC) racemose, protrudes upper edge of poreplate (PPl); posterior margin of triangular posterior stalk (PS) almost straight; lateral margins of genital atrium (GA) slightly sclerotized (Fig. 8).

Variation.

Males (N = 14): BL 12.64–16.98, CL 6.27–7.56, CW 6.02–7.56, OL 6.95–8.50, OW 4.60–6.40; females (N = 5): BL 10.55–14.24, CL 5.26–6.37, CW 4.47–5.63, OL 5.24–7.12, OW 4.33–4.96. The number of denticles on the promargin of male cheliceral groove varies from 10 to 13 (N = 12); in females, the number of denticles on the promargin of cheliceral groove varies from 11 to 12 (N = 5). The number of setae on tibial apophysis varies from 4 to 6.

Etymology.

The species epithet “ nawngau ” refers to the type locality, Nawng Au Village; it is treated as a noun in apposition.

Distribution.

Shan State, (Kyaukme District), Myanmar.