Anthomyia acklandi Vikhrev 2022, sp. nov.

Anthomyia acklandi sp. nov.

https://zoobank.org/NomenclaturalActs/E20F2078-

CDE5-475C-8F6D-472602F3E828 Figs. 1–5

Holotype ♂: INDONESIA, Papua prov., Lake Habbema, 3350 m, 4.14 ° S, 138.71 ° E, 19– 26.12.2014, N. Vikhrev.

Paratypes: the same locality, 19– 26.12.2014, N. Vikhrev. 13♂, 6♀; 10– 12.12.2017, N. Vikhrev, 3♂ 3♀. The entire type series is deposited in Zoological Museum of Moscow University (= ZMUM).

Description. Head. Eyes bare, distance between eye margins at narrowest part slightly narrower than width of postpedicel (Fig. 2). Fronto-orbital plates at posterior 2/3 black and narrow, at anterior 1/3 grey and widened, with 3 pairs of inclinate setae on anterior 1/3. Frons narrow, yellowish-grey, with rather strong crossed interfrontals. Parafacials grey, with dark spots at upper and lower thirds. Mouthedge slightly protruding. Cheek about 1.5x as wide as width of postpedicel. Postpedicel 2.25x as long as wide. Longest aristal hairs slightly longer than basal diameter of arista. Proboscis elongated (Fig. 1), about 2/3 as long as fore tibia.

Thorax densely grey dusted over dark ground-colour; with contrasting black pattern on dorsal surface. Presutural area of scutum with a pair of rectangular black spots merged in front and diverging posteriorly (Fig. 2); postsutural area with black transverse band from suture to almost level of 3 rd post dc (Figs. 1, 2). Scutellum black with only extreme apex pale, with setulae on ventral side. Black colour is not extended on pleura (darkening of anepisternum on Fig. 1 is a result of reflection). 2 + 3 dc; 3 pairs of presutural ac in rows separated by more than distance from each to adjacent dorsocentral row, without additional setulae in between. Prealar seta as strong as posterior notopleural; katepisternals 2 + 2, lower posterior weak; proepisternum with 5–6 fine setulae. Wing with costal basal marginal spinules strong; anterior margin of wing darkened.

Legs dark. t1 with 1 pv and 1 ad; t2 with 1 ad, 2 pd and 1 pv; t3 with 1 av, 3 ad, 2 pd, 4–5 fine short pv at middle.

Abdomen grey dusted, tergites 1 + 2 to 5 with tridental black marks, less distinct on tergite 1 + 2 (Fig. 3). Surstyli elongated, widened in middle, with pointed apex; without stronger inner setae, cercal plate without specialised flattened apical setulae (Fig. 4); sternite 5 with a pair of strong apical setae (Fig. 5).

Female similar to male except for wide frons (40% of head width) and shape of abdomen, which is pointed at the tip.

Diagnosis. A. acklandi sp. nov. has thorax with typical “ Anthomyia -pattern” and setulose proepisternum. It clearly differs from all Asian Anthomyia by presutural area of scutum with a pair of rectangular black spots merged in front and diverging posteriorly. Male has widened frons and simplified structure of genitalia.

Etymology. Named in honour of Michael Ackland, foremost expert on the family Anthomyiidae, who died in 2021.

Anthomyia fumipennis Stein, 1919 Anthomyia fumipennis Stein, 1919: 210–211 Figs. 6–10

Material examined: INDONESIA, Papua Prov., Wamena env., Baliem Resort, 2000 m, 4.06 ° S, 139.03 ° E, 16– 25.12.2014, N. Vikhrev, 5♂, 2♀; 11– 16.12.2017, N. Vikhrev, 10♂, 1♀; 14 km W of Wamena, 2800 m, 4.137 ° S, 138.826 ° E, 7.12.2017, N. Vikhrev, 1♂, 3♀ (all ZMUM).

Description of male. Head. Eyes bare, eye margins separated on diameter of ocellus (Fig. 7). Fronto-orbital plates linear, black and narrow at posterior half, at anterior half widened and with 5 pairs of inclinate setae and minute crossed interfrontal setulae. Parafacials grey, with dark spots at upper and lower thirds. Mouthedge not protruding. Cheek about 1.5x as wide as width of postpedicel. Postpedicel 3x as long as wide. Longest aristal hairs half as long as width of postpedicel. Proboscis not elongated.

Thorax densely grey dusted over dark ground-colour; with contrasting black pattern on dorsal surface. Presutural area of scutum with black transverse band from neck to level of 2 nd prst dc (Fig. 7); postsutural area with black transverse band from suture to level of 3 rd post dc; pre- and postsutural bands narrow merged at level of intraalar setae (Fig. 7). Black colour also extended on pleura: most anepisternum and anterior part of katepisternum (Fig. 6). Scutellum black, with setulae on ventral side. Prealar seta as strong as posterior notopleural; katepisternals 2 + 2, proepisternum with 9–12 setulae, lower posterior weak. 2 + 3 dc; 1 strong pair of presutural ac without additional setulae in between. Wing with marginal spinules short; whole wing surface darkened.

Legs dark. t1 with 1 pv; t2 with 1 ad; 2 pd and 1 pv; t3 with 1 av, 3 ad, 2 pd and 4–5 short pv at middle.

Abdomen grey dusted, tergites 1+2 to 5 with tridental black marks, more extended than in A. acklandi sp. nov. (Fig. 8). Surstyli simple, short, with blunt apex (Fig. 9), without stronger inner setae; cercal plate without specialised flattened apical setulae. Sternite 5 without strong setae (Fig. 10).

Female similar to male except for wide frons (40% of head width) and shape of abdomen, which is pointed at the tip. Dark area on pleura less extensive.

Distribution. Central regions of New Guinea at altitudes 1700–2800 m.