Giraudia japonica sp. nov.

(Standard Japanese name:

Kimon-kuro-chibi-togari-himebachi)

(Figs. 1A, G, 3A, B, 4A–E)

Holotype. KPM-NK 69946, F, JAPAN, Tochigi Pref., Nasushiobara City, Amayu - Shiobara, 12. X. 2009, E. Katayama leg.

Paratype. KPM-NK 69947, F, JAPAN, Nagano Pref., Karuizawa, 8. VIII. 1952, R. Ishikawa leg.

Description. Female (n=2). Body length 9.5–10.5 (HT: 9.5) mm, mat, covered with golden setae.

Head 0.6 times as long as wide. Clypeus transverse (Fig. 1A), 0.3 times as long as wide, punctate dorsal 0.6 and almost smooth ventral 0.4, covered with long setae (Fig. 4B), lower margin flattened. Face 0.3–0.35 (HT: 0.35) times as long as wide, sparsely punctate, convex medially (Fig. 4B). Frons slightly concave, entirely mat except for narrow smooth area above antennal sockets. Occipital carina complete, its lower end joined with hypostomal carina. Lower tooth of mandible distinctly longer than upper tooth (Fig. 1A). MSL 0.7–0.8 (HT: 0.7) times as long as BWM. OOL 1.1–1.2 (HT: 1.2) times as long as POL. Antenna with 32–33 (HT: 32) flagellomeres. F1 0.8 times as long as F2. Apical half of flagellum flattened (Fig. 4A), distinctly wider than basal part.

Mesosoma. Epomia present, weak and short. Lateral area of pronotum partly weakly rugose, without longitudinal striation. Notauli distinct on anterior 0.33 of mesoscutum. Scutellum weakly convex. Epicnemial carina distinct, its dorsal end not joined with anterior margin of mesopleuron and subalar prominence. Mesopleuron and metapleuron finely punctate. Pleural carina of propodeum complete. Lateral longitudinal carina, lateromedian longitudinal carina and posterior transverse carina largely indistinct. Other carinae absent. Area superomedia indistinctly defined (Fig. 4C). Propodeal apophysis obtusely present. Propodeal spiracle elongate. Fore wing length 9.0–9.5 (HT: 9.0) mm. Areolet present, hexagonal, with sides convergent anteriorly (Figs. 1G, 4A). Nervulus opposite to basal vein (Fig. 4A). Nervellus intercepted posterior to midpoint (Fig. 4A). Tibiae with some long setae (Fig. 4D). Hind femur 4.5–4.8 (HT: 4.8) times as long as maximum depth in lateral view. Hind TS2: TS1, TS3, TS4 and TS5 = 2.0: 3.7–4.0 (HT: 3.7), 1.4–1.6 (HT: 1.4), 0.8–1.0 (HT: 0.8) and 1.3–1.5 (HT: 1.3).

Metasoma. T1 2.3–2.7 (HT: 2.3) times as long as maximum width, without dorsolateral and lateromedian longitudinal carinae, its postpetiole covered with fine and distinct sculptures (Fig. 4E). T2 0.65–0.7 (HT: 0.7) times as long as maximum width. Ovipositor sheath 1.25 times as long as hind tibia. Apex of ovipositor as Figs. 3A and B.

Coloration (Figs. 4A–E). Body (excluding wings and legs) black, except for: dorsal part of facial orbit, frontal orbit, vertical orbit, palpi, median part of anterior margin of collar, median small area of T1 and T2, narrow area of posterior margin of T2 and T3 and ovipositor reddish brown; F6–F11 (sometimes also F5 and dorsal part of F12), scutellum and postscutellum yellow. Wings hyaline, except for: stigma yellowish brown; veins brown except for yellow wing base. Legs black, except for: apex of femora, tibiae, tibial spurs and tarsi reddish yellow.

Male. Unknown.

Distribution. Japan (Honshu).

Remarks. This species resembles G. gyratoria (Thunberg, 1822) in the yellow scutellum and the normal shaped wings, but can be distinguished by the following combination of character states: postpetiole covered with fine and distinct sculptures (Fig. 4E) (smooth and without sculpture in G. gyratoria); base of antenna black (Fig. 4A) (with conspicuous reddish brown area in G. gyratoria): hind trochanter and trochantellus blackish brown (Fig. 4A) (red in G. gyratoria); metasomal tergites more or less darkened (Fig. 4A) (with a conspicuous large red area in G. gyratoria). This species can be easily distinguished from other Japanese species, G. spinosa Uchida, 1936, and G. teranishii Uchida, 1930, by the yellow scutellum (Fig. 4A) (black in other Japanese species).