Eupyrochroa insignita

Eupyrochroa insignita (Fairmaire, 1894)

Pyrochroa insignita Fairmaire, 1894: 220 (Type locality: “Se-Pin-Lou-Chan, Thibet” [located in Ya’an, Sichuan, China; see Qiu et al. 2013]. Type deposition: MNHN).

Pyrochroa insignita var. limbaticollis Pic, 1909: 330 (Type locality: “ Chine ”. Type deposition: MNHN).

Eupyrochroa insignita: Blair, 1914: 316; Wu, 1937: 423; Hua, 2002: 131; Pollock & Young, 2008: 415; Young et al., 2020: 566; Pan et al., 2021: 17.

Eupyrochroa limbaticollis: Blair, 1914: 316; Kôno, 1935: 160; Wu, 1937: 423; Hua, 2002: 131; Young, 2004: 195; Pollock & Young, 2008: 415; Young et al., 2020: 566; Pan et al., 2021: 17. syn. nov.

Eupyrochroa insignita var. rufoverticalis Pic, 1915: 4 (Type locality: “Chine méridionale: Tali ” [Dali, Yunnan, China]. Type deposition: MNHN).

Pseudopyrochroa limbaticollis var. immaculata Pic, 1923: 19 (Type locality: “ Chine ”. Type deposition: MNHN).

Pseudopyrochroa immaculata: Hua, 2002: 131.

Redescription. Adult. Male (Fig. 3A): Body length 18.0–25.0 mm, humeral width 3.5–6.0 mm. Body generally black, except elytra burnt orange to crimson, pronotum black to reddish black (Fig. 1), scutellum black, tan to red (Fig. 1), mandibular apices and anterior margin of clypeus reddish brown. In some individuals, head (including antennae), femora, and tibiae uneven in color, dark red to reddish black. Body densely covered with yellow-brown semi-erect short setae, but setation tawny on dorsal surface of head and black portion of pronotum, and orange-red on red portion of pronotum and elytra.

Head (Fig. 3B) with small, moderately dense punctures, diameter of punctures less than spacing between punctures, each puncture unisetose. Frons slightly convex, with a transverse depression across interocular surface; cranial pit large, shallow, scutiform, with dense yellow-brown setation inside, edges nitid and complete. Vertex smooth, slightly convex. Compound eyes moderate, widely spaced (OI=44.2–53.2). Genae strongly angulate. Antennae (Fig. 3D) longer than half length of elytra; scape long and stout, enlarged distally, nearly subequal to length of flagellomere I; pedicel shortest; flagellomeres I–III gradually shortened, III–VI similar in length, VII–IX gradually shortened, rami of III–X gradually elongated, terminal flagellomere longest.

Pronotum (Figs. 1A–C; 3A) nearly rectangular, with angular sub-basal projections, wider than long (PL/PW: 0.56–0.81); punctures dense, unisetose; disc with a longitudinal, mesal furrow and transverse depression on each side of meson. Scutellum longer than wide. Each elytron with three distinct longitudinal ridges. Tibiae slender, tibial spurs short. Prothoracic tarsomere V longest, I second, II–IV subequal in length; mesothoracic tarsomeres I and V similar in length, II–IV slightly shorter, subequal in length; metathoracic tarsomere I longest, followed by IV, II and III gradually shorter. Pretarsal claws simple.

Posterior margins of abdominal sternites III–VI subparallel, that of VII obtusely and shallowly emarginate and VIII acutely V-shaped, emarginate. Parameres approximately as long as phallobase in lateral view (Fig. 3G), fused along basal 9/10 and depressed around inner margins of apices in dorsal view (Fig. 3F); penis (Fig. 3H, I) tapered, broadest in middle, apex pointed in dorsal view and enlarged in lateral view, with short, paired basal struts.

Female: Body length 17.0–24.0 mm, humeral width 3.5–6.0 mm. Head (Fig. 3C) without cranial pit; antennal rami distinctly shorter than those of males (Fig. 3E); OI (40.4–55.3) with wider range than males. Posterior margins of abdominal sternites VII–VIII almost straight. Other features largely as in males.

Larva. Mature larvae (Fig. 4A) attain lengths (mesal labral apex to apices of urogomphi) of 26.0–41.0 mm and widths (across widest portion of abdominal segment VIII) of 4.0–6.0 mm. Body orthosomatic with sides subparallel, amber in color, moderately sclerotized, mandibles and urogomphal plate heavily sclerotized, orange-brown to dark brown; overall surface with very few scattered long yellow setae; thoracic and abdominal terga each with a weak parabasal ridge.

Head (Fig. 4B–C) oval, prognathous, mouthparts retracted but with mandibles and maxillary apices protruding. Epicranial suture lyriform with stem short, frontal arms extending anteriolaterally to bases of antennal insertions. Four stemmata on each side, around antennal insertions, symmetrical arrangement. Antennal insertions fully exposed, each antenna 3-segmented, filiform; antennomere I 0.05–0.08 longer than II and III, tapering; I constricted at mid-length, I and II slightly enlarged to apex; III thinnest, apex sharp. Labrum semicircular, slightly acuminate anteriomesally. Mandibles well sclerotized, asymmetrical, mola of each mandible well developed, left mandible bearing a prominent molar tooth and three apical teeth, right mandible with two apical teeth. Maxillae each with 1-segmented cardo which is diagonally folded upward upon itself toward stipes and thus appearing 2-segmented; a well-developed, undivided, pad-like maxillary articulating area; ventral surface of stipes bearing few scattered spiniform setae; each maxillary palpus 3-segmented, filiform, palpomere I stubby and tapering, I–II with short thick yellow setae. Mentum inverted trapezoidal; submentum rectangular, with a pair of setae at center; ligula well developed, with sparse short setae; each labial palpus 2-segmented, palpomere I 1.65× as long as II and stronger than II. Hypostomal rods well developed, divergent, extension lines form right angles, subparallel to gular suture.

Thorax (Fig. 4A) flattened, well developed, 3-segmented. Prothorax (Fig. 4B, C) subrectangular. Meso- and metathorax arc-shaped on both sides, nearly teardrop-shaped. Legs (Fig. 4A–C) 5-segmented, vestiture consisting of sparse, short setae.

Abdomen (Fig. 4A) with 10 relatively flat segments, segments I–VIII with curved lateral margins, segments I–VII subequal in length and width, VIII longest and twice of VII in length, sides round (Fig. 5). In ventral view, VIII (Fig. 5B) with a well developed, inverted U-shaped posterior emargination and a narrow, strongly acuminte, apical tooth-like process bilaterally. Tergite IX (Fig. 5) strongly sclerotized, with acutely rounded lateral lobes extending forward on base of lateral margin; anterior margin U-shaped, concave; surface of urogomphal plate bearing numerous well-developed callosities, several setigerous calli, and sparse, long setae at edges; in particular, one pair of calli on dorsolateral surface and another pair at base of urogomphi; urogomphi elongate, straight, slightly divergent, tapering and acuminate apically; ventral surface of urogomphal plate depressed at center, forming a narrowing longitudinal groove caudally. Urogomphal plate possessing a heavily sclerotized, inverted sub-triangulate urogomphal lip (Figs. 4D, 5), ventral surface smooth, apex tapering; two deep urogomphal pits (Fig. 4D) between urogomphi, with apparent parallel rugulae. Anus (Fig. 5B) positioned at junction of sternites IX and X. Sternite X with uneven rectangular edges.

Paired spiracles oval, located at ventrolateral margin along anterior end of mesothoracic laterotergite, dorsolateral margin of abdominal tergite I, ventrolateral margins of abdominal laterotergites II–VII, and distal about 1/3 (0.32) of length of laterotergite VIII ventrolaterally (Fig. 5B).

Type material examined. Eupyrochroa insignita var. rufoverticalis: 1 ex., labelled “ Tali, Haut Yunnan ”, “notabilis, Blair (ex expl. communique)”, “type”, “TYPE”, “v. rufoverticalis Pic ” (MNHN). Pyrochroa insignita var. limbaticollis: 1 ex., labelled “Kuatun”, “ Chine ”, “ Pyrochroa insignita Fairm. ”, “type”, “ TYPE ”, “ insignita var. nov. limbaticollis Pic ” (MNHN). Pseudopyrochroa limbaticollis var. immaculata: 1 ex., labelled “Hoachou, 15.5.18”, “ type ”, “v. immaculata Pic ” (MNHN).

Other materials. Adults: CHINA: 1♀, Chine (MNHN). Henan: 1♀, 2021. VI.8, Henan, Nanyang, Neixiang, Baotianman, Hui Guo leg., elev. 1450 m, Museum of Hebei University (MHBU); 1♀, 2022. V.27, Henan, Xinyang, Shangcheng County, Changzhuyuan, Huangbaishan, Hui Guo leg., Museum of Hebei University (MHBUa). Shaanxi: 1♀, 2019.VII.6–15, Shaanxi, Yan’an, Fu County, Ziwuling Reserve, Shihuigou, Museum of Hebei University; [2 nd label]: P1A1 (MHBUa); 1♀, 2021. V.25, Shaanxi, Baoji, Chencang District, Mt. Jiulong, 34°20'31.95"N 106°52'10.83"E, elev. 811 m, Juan Li et al. leg., Museum of Hebei University; [2 nd label]: P1H4 (MHBUa); 1♂, 2021. Ⅴ.26, Shaanxi, Baoji, Chencang District, Mt. Jiulong, Juan Li et al. leg., Museum of Hebei University; [2 nd label]: P1G8 (MHBUa); 2♀♀, 2021. V.28, Shaanxi, Baoji, Long County, Badu Town, 34°41'18.63"N 106°47'9.14"E, elev. 1226 m, Juan Li et al. leg., Museum of Hebei University; [2 nd label]: P1G9/P1H3 (MHBUa); 1♀, 2021. Ⅴ.30, Shaanxi, Baoji, Feng County, Tangzang Town, Juan Li et al. leg., Museum of Hebei University; [2 nd label]: P1G10 (MHBUa); 1♂, 2019. Ⅴ.29, Shaanxi, Baoji, Mei County, Mt. Taibai, Fengshan Gully, Yu-Chen Zhao leg., Museum of Hebei University; [2 nd label]: 34.1275°N 107.9010°E, elev. 750 m, Museum of Hebei University (MHBU); 1♀, Shaanxi Prov., Taibai Shan above Houshenzi, 1300–1700 m, 9. VI.–3.VII.1998, leg. P. JÄGER & J. MARTENS (SMNS); 1♀, SHAANXI, Taibaishan Mts., env. Haozhanzi, 1500–2000 m 33:52’N 107:44E, VI – VII.2000 A. Plutenko leg. (DYCC); 1♀, CHINA: SHAANXI, Taibaishan Mts., env. Haozhanzi, 1500–2000 m 33:52’N 107:44E, VI –VII.2000 A. Plutenko leg. (SKCC). Gansu: 1♀, 2021. Ⅴ.13, Gansu, Longnan, Hui County, Yanping Forestry Farm, Juan Li et al. leg., Museum of Hebei University; [2 nd label]: P1H1 (MHBUa); 1♀, 2021. Ⅴ.18, Gansu, Longnan, Hui County, Gaoqiao Forestry Farm, Juan Li et al. leg., Museum of Hebei University; [2 nd label]: P1H2 (MHBUa); 1♀, No. 0054, 1992. VI.5, Gaoqiao, Guo-Qing Song leg. (MHBU); 2♀♀, 2021. VI.1, Gansu, Longnan, Hui County, Jiangluo, Liujiaba, 33°55'48.84"N 105°52'10.18"E, elev. 1203 m, Qi Liu & Rui Liu leg., Museum of Hebei University; [2 nd label]: P1H8/P1H9 (MHBUa); 2♂♂, 2♀♀, 2021. VI.03, Gansu, Hui County, Jiangluo, Rui Liu leg., Museum of Hebei University; [2 nd label]: 33°55′1.27″N 105°58′52.89″E, elev. 1187 m, Museum of Hebei University (MHBU); 3♀♀, 2021. VI.02, Gansu, Hui County, Mayan, Baiya, Qi Liu leg., Museum of Hebei University; [2 nd label]: 34°3′38.11″N 105°45′12.15″E, elev. 1357 m, Museum of Hebei University (MHBU); 1♂, 2021. VI.02, Gansu, Hui County, Mayan, Baiya, Qi Liu leg., Museum of Hebei University; [2 nd label]: 34°3′38.11″N 105°45′12.15″E, elev. 1357 m, Museum of Hebei University; [2 nd label]: P1 (MHBU); 1♀, Mayan, 1989. VI.6 (MHBU); 2♂♂, 1♀, 2021. Ⅴ.27, Gansu, Liangdang, Zhangjiazhaiziling, Qi Liu leg., Museum of Hebei University; [2 nd label]: 34°12′15.66″N 106°28′0.65″E, elev. 1802 m, Museum of Hebei University (MHBU); 1♀, 2021. VI.4, Gansu, Longnan, Liangdang County, Yangdian, Luojiazhuang, 33°54'6.54"N 106°23'49.65"E, elev. 1183 m, Qi Liu & Rui Liu leg., Museum of Hebei University; [2 nd label]: P1H10 (MHBUa); 1♂, No. 84, 1992. VI.21, Majiaba, Xiao-Li Ma leg. (MHBU); 1♀, No. 363, 1992.VII.29, Xinglong, Jin-Ping Zhang & Jin-Yong Zhao leg. (MHBU); 2♀♀, Gansu, Xiaolongshan, 1996. VI.24 (MHBU); 2♀♀, Gansu, Xiaolongshan, Museum of Hebei University (MHBU); 1♀, No. 105, Longmen (MHBU); 1♀, 2021. V.21, Gansu, Maiji, Guanyin Forestry Farm, Juan Li et al. leg., Museum of Hebei University; [2 nd label]: P2C2 (MHBUa); 1♀, 2021. V.29, Gansu, Tianshui, Qinzhou District, Niangniangba, Hekou, 34°15'40.12"N 105°53'52.14"E, elev. 1408 m, Qi Liu & Rui Liu leg., Museum of Hebei University; [2 nd label]: P1H5 (MHBUa); 1♂, 1♀, 2021.VII.7, Gansu, Tianshui, Shanmen, 34°41'30.24"N 106°21'37.41"E, elev. 1600 m, Qi Liu & Quan-Yu Ji leg., Museum of Hebei University; [2 nd label]: P1H6/P1H7 (MHBUa); 1♀, 2009.VII.11–12, Gansu, Zhouqu County, Guazigou, Zhen-Hua Gao & Yi-Ping Niu leg., Museum of Hebei University (MHBU); 1♂, S GANSU, Tsagan (70 km W Wudu), 2300–2400 m, 16–17/vi/2001, S. Kazantsev leg. (SKCC); 2♂♂, S GANSU, Tsagan (70 km W Wudu), 2300–2400 m, 16–17/vi/2001, S. Kazantsev leg. (DYCC); 1♀, Hweisin, Kansu [= Gansu]; [2 nd label, handwritten on old, green paper]: 10338; [3 rd label]: ex coll. Witold Eichler, [not legible], Inst. Zool. P.A.N., Warszawa, 64163 (IZCW). Jiangsu: 1♂, Nanjing, Mt. Zijin, 2006.IV.27, Zhen Wang leg. (MHBU); 2♀♀, Lungtan bei Nanking; Prov. Kiangsu; 13.5.1933. H. Höne. (ZMHB); 1♀, Lungtan bei Nanking; Prov. Kiangsu; 13.5.1933. H. Höne. (DYCC). Zhejiang: 1♀, Tsche-kiang; Wen chow [= Wenzhou] (ZMHB); 1♀, (Zhejiang) Tianmu Shan, pass 25 km, NNW Linan, 620–820 m, 30˚25’40”N 119˚35’30”E (creek valley, bamboo, mixed forest, beaten from vegetation), 16. VI.2007 D. W. Wrase [37A] (NKME). Hubei: 2♂♂, 2♀♀, Hubei Prov., alt. 440 m, Nanmuhe, Wufeng Pref., 27.IV.2004, N. Ohbayashi leg. (DYCC); 5♂♂, 9♀♀, Hubei Province; Mt. Wangfoshan; Alt. 1940 m; Baokang County; 20–28 June 2008; purchase ex: Li Jingke (DYCC). Hunan: 2 exx., Hunan, Baojing, Yezhuping, Huixizu (elev. 520 m), 2018.IV.17, Kai-Qin Li leg. (KNHM); 2 exx., Hunan, Wulingyuan, Yixiantian (elev. 430 m), 2018.IV.18, Kai-Qin Li leg. (KNHM); 1 ex., Hunan, Wulingyuan, Tiechang, reserve of giant salamander (elev. 300 m), 2018.IV.19, Kai-Qin Li leg. (KNHM). Fujian: 1♂, Kuatun, 7.5.46, Tschung Sen (MNHN); 1♀, KUATUN, KUKIEN 9.5.46, (TSCHUNG SEN.); [2 nd label] Sammlung, J. KLAPPERICH, SMNS 1990; [3 rd label][handwritten - perhaps Pic’s hand — not legible]; [4 th label] [likely Pic’s hand] Eupyrochroa limbaticollis Pic (SMNS); 2♀♀, KUATUN, KUKIEN, 12.5.46 (TSCHUNG SEN.) (SMNS); 2♀♀, KUATUN, KUKIEN, China 12.5.46 (TSCHUNG SEN.) (DYCC); 1♀, KUATUN, KUKIEN, China 12.4.46 (TSCHUNG SEN.) (DYCC); 2♂♂, 3♀♀, DALINGSHAN, JIANLING FUJIAN, April 2003; Elev 1600 m, leg: LI (DYCC); 2♂♂, 7♀♀, Fukien; Shaowu, Tachuland; 6–10 May 1943; T. C. Maa leg. (BPBM); 2♂♂, 2♀♀, Fukien; Shaowu, Tachuland; 6–10 May 1943; T. C. Maa leg. (DYCC); 1♂, 1♀, Fukien; Shaowu, Tachuland; 12–13 May 1943; T. C. Maa leg. (BPBM); 1♂, 1♀, Fukien; Shaowu, Tachuland; 12–13 May 1943; T. C. Maa leg. (DYCC); 2♂♂, 3♀♀, Fukien: Shao Wu: Ta Chu Lan; 27.iv.1942; [2 nd label]: T. C. Maa, Collector (CASC); 1♂, 1♀, Fukien: Shao Wu: Ta Chu Lan; 27.iv.1942; [2 nd label]: T. C. Maa, Collector (DYCC); 2♀♀, Fukien Prov.; Shaowu: Tachulan; 3.v.1942; [2 nd label]: T. C. Maa, Coll. (CASC); 1♂, FUKIEN, S. China: Shaowu, Tachuland; 7.iv.1942; [2 nd label]: T. C. Maa, Collector (CASC); 1♀, FUKIEN, S. China: Shaowu, Tachuland; 10.vi.1942; [2 nd label]: T. C. Maa, Collector (CASC); 2♀♀, FUKIEN, S. China: Shaowu, Tachuland; 12.v.1942; [2 nd label]: T. C. Maa, Collector (CASC); 1♀, FUKIEN, S. China: Shaowu, Tachuland; 12.v.1942; [2 nd label]: T. C. Maa, Collector (DYCC); 1♂, FUKIEN, S. China: Shaowu, Tachuland; 15.iv.1942; [2 nd label]: T. C. Maa, Collector (CASC); 1♂, 2♀♀, FUKIEN, S. China: Shaowu, Tachuland; 19.iv.1942; [2 nd label]: T. C. Maa, Collector (CASC); 1♂, FUKIEN, S. China: Shaowu, Tachuland; 19.iv.1942; [2 nd label]: T. C. Maa, Collector (DYCC); 1♂, FUKIEN, S. China: Shaowu, Tachuland; 22.iv.1943; [2 nd label]: T. C. Maa, Collector, BISHOP Mus. (CASC); 4♂♂, 4♀♀, FUKIEN, S. China: Shaowu, Tachuland; 21.iv. 1942; [2 nd label]: T. C. Maa, Collector (CASC); 2♂♂, 2♀♀, FUKIEN, S. China: Shaowu, Tachuland; 21.iv.1942; [2 nd label]: T. C. Maa, Collector (DYCC); 2♂♂, 2♀♀, FUKIEN, S. China: Shaowu, Tachuland; 26.iv.1942; [2 nd label]: T. C. Maa, Collector (CASC); 1♂, 3♀♀, FUKIEN, S. China: Shaowu, Tachuland; 27.iv.1942; [2 nd label]: T. C. Maa, Collector, BISHOP Mus. (CASC); 1♂, 1♀, FUKIEN, S. China: Shaowu, Tachuland; 27.iv.1942; [2 nd label]: T. C. Maa, Collector, BISHOP Mus. (DYCC); 1♂, 8♀♀, FUKIEN, S. China: Shaowu, Tachuland; 30.iv.1942; [2 nd label]: T. C. Maa, Collector, BISHOP Mus. (CASC); 2♀♀, FUKIEN, S. China: Shaowu, Tachuland; 30.iv.1942; [2 nd label]: T. C. Maa, Collector, BISHOP Mus. (DYCC); 1♂, FUKIEN, S. China: Shaowu, Tachuland; 2.v.1943; [2 nd label]: T. C. Maa, Collector (CASC); 1♂, FUKIEN, S. China: Chungan: Tsilichi, -ao to Sanchiang, 194 T. Maa; [2 nd label]: 28.iv.1945 (CASC); 1♂, 1♀, FUKIEN, S. China: Chungan: Tsilichi, -ao to Sanchiang, T. Maa; [2 nd label]: 29.iv.1942 (CASC); 1♂, FUKIEN, S. China: Chungan: Tsilichi, -ao to Sanchiang, T. Maa; [2 nd label]: 29.iv.1942 (DYCC). Sichuan: 1♀, Siào-Lòu, Chasseurs indigènes, 1894 (MNHN); 1♀, 2022. V.19, Sichuan, Qionglai, Mt. Tiantai, Xiaojiawan, 103°08'05"N 30°17'05"E, elev. 862.22 m, Zi-Yuan Hu leg., Museum of Hebei University (MHBUa); 1♂, 1♀, Su-Tchuen, Ya-Tchéou, 1897 (MNHN); 2♀♀, Su-Tchuen, Chass. Thibétains, 1897 (MNHN); 1♀, Mou-Pin, 1897, ex R P Déjean (MNHN); 1♀, North Sichuan, 60km. E. of Songpan env., Minshan Mts. 1000 m, 1–10.vii.2004, V. Patrikeev; [2 nd label - reversed] BMNH {E}, 2005-104, MVL Barclay (BMNH); 3♀♀, N. Sichuan, Minshan Mountains, 15 km S. Nanping; 2000 m, 20 June 2005, V. Patrikeev leg., Exchange with Ilya Osipov (DYCC); 1♂, 1♀, Prov. Sichuan, Wo Long Gebiet, 1500 m, 2. V.2001, MENRAD leg. (SMNS); 1♀, N. Sichuan, Valley NNW MUZUO, 32.40–44N 104.32–30E, 2000–2700 m, J Kaláb leg., 28.v– 6.vi.2006; [2 nd label - reversed] BMNH {E}, 2006-123, MVL Barclay; [3 rd label]: Retained by Permission, from series located at, BMNH, London, 29 November 2010 (DYCC); 1♀, N. Sichuan, Valley NNW MUZUO, 32.40–44N 104.32–30E, 2000–2700 m. J. Kaláb leg., 28.v–6.vi.2006 (BMNH); 2♀♀, SICHUAN, Nanjiang, 18.5.– 22.5.2010, lgt. E. Kučera (BMNH). Yunnan: 1♀, CASENT, 6005452; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang,; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (CASC); 1♀, CASENT, 6005453; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (CASC); 1♀, CASENT, 6005454; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (CASC); 1♀, CASENT, 6005455; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (CASC); 1♀, CASENT, 6005456; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (CASC); 1♀, CASENT, 6005457; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (DYCC); 1♀, CASENT, 6005458; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (DYCC); 1♀, CASENT, 6005459; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao, headwaters Longchuan Jiang; [3 rd label]: N25.73947° E098.69630°, 2010 m, 16 May 2006, Stop #2006-025A, D. H. Kavanaugh & R. L. Brett collectors (DYCC); 1♀, CASENT, 6005513; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, 0.4 km N of Dahetou Lingganjiao at Longtang He; [3 rd label]: N25.74277° E098.69691°, 2020 m, 17 May 2006, D. H. Kavanaugh & R. L. Brett collectors, Stop #2006-027 (CASC); 1♀, CASENT, 6005516; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, 0.4 km N of Dahetou Lingganjiao at Longtang He; [3 rd label]: N25.74277° E098.69691°, 2020 m, 17 May 2006, D. H. Kavanaugh & R. L. Brett collectors, Stop #2006-027 (DYCC); 1♀, [CHINA]: CASENT, 6005637; [2 nd label]: CHINA, Yunnan, Tangchong County, Jietou Township, Dahetou Lingganjiao; [3 rd label]: N25.73947° E098.69630°, 2010 m, 20 May 2006, Stop #2006-035, D. H. Kavanaugh & R. L. Brett collectors (CASC). Xizang: 1♀, 2021.VII.19, Xizang, Zayü County, Chawalong, Gunshi, 28.566974°N 98.308763°E, elev. 2396.12 m, Guo-Dong Ren & He Shi leg., Museum of Hebei University; [2 nd label]: P1I1 (MHBUa). LAOS: Houaphanh: 1♀, NE LAOS, Hua Phan prov. [Houaphanh], Ban Saleui Plu Pan (Mt.), 1300–1900 m, 27.IV.-1. VI. 2011, 20˚12′N, 104˚01′E, leg. C. Holzschuh; [2 nd label]: BMNH{E}, 2012–14, C. Holzschuh (BMNH).

Larvae: CHINA: Gansu: 10 exx., 2020.VIII.3, Gansu, Maiji, Tailu Forestry Farm, 34°27.489'N 106°13.014'E, elev. 1451 m, Guo-Dong Ren et al. leg., Museum of Hebei University; [2 nd label]: P1E5/P1E7/P1E8/P1E9 (MHBUa); 6 exx., 2020.VIII.3, Gansu, Maiji, Tailu Forestry Farm, 34°30.641'N 106°13.248'E, elev. 1138 m, Guo-Dong Ren et al. leg., Museum of Hebei University; [2 nd label]: P1F7–P1F10 (MHBUa).

Distribution. China: Henan, Shaanxi, Gansu, Jiangsu, Zhejiang, Hubei, Hunan, Jiangxi, Fujian, Taiwan, Sichuan, Yunnan, Xizang; Laos: Houaphanh.