Landouria madurensis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.

Landouria madurensis sp. nov.

urn:lsid:zoobank.org:act: 024A8B78-91C2-49CA-8721-C0B8CCC46982

Figs 8, 22, 36, 53, 57; Tables 3–4

Helix squamulosa Martens, 1867: 266, pl. 13, fig. 10 (“ Insel Madura (an der Nordostseite von Java), bei Sumanap”) (objective junior synonym of Helix squamulosa Mousson, 1856). Syn. nov.

Helix squamulosa – Pfeiffer 1871 in 1870–1876: pl. 117, figs 13–15; 1872 in 1870–1876: 41. Landouria squamulosa – van Benthem Jutting 1941: 311.

Diagnosis

Landouria madurensis sp. nov. is characterized by a sharply keeled shell with longish scales all over the teleoconch, a wide umbilicus, a long penis, which is divided into a longer, broader distal and a shorter, narrower proximal part, and a long flagellum, which gradually tapers towards its proximal end.

Etymology

Landouria madurensis sp. nov. is named after its occurrence on Madura Island.

Material examined

Holotype

INDONESIA • holotype (det. anat.); East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest; 6°54′01″ S, 113°12′08″ E; 17 m a.s.l.; 14 May 2014; A.S. Nurinsiyah and T. Setiadi leg.; D = 11.4 mm, H = 6.5 mm; MZB 21432.

Paratypes

INDONESIA • 1 spec.; same data as for holotype; MZB 19209 • 4 spec.; same data as for holotype; MZB 19210 • 1 spec.; same data as for holotype; ZMH 133408 • 4 spec.; same data as for holotype; ZMH 133437 • 2 spec. (also syntypes of H. squamulosa); Sumenep (see Martens 1867: 267); 7°01′ S, 113°52′ E; ZMB 7155.

Description

SHELL (Figs 8, 57; Tables 3–4). Depressed conical, with 5.5–5.75 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; without distinct incised spiral lines; longish scales all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, very wide, comprising 38–40% of shell diameter, hardly obscured by columellar edge.

GENITALIA (Figs 22, 36; Table 4). Atrium short; penis long, divided into a longer, broader distal and a shorter, narrower proximal part, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to middle part of cylindrical epiphallus; flagellum long, broadest in its middle part, gradually tapering towards its proximal end, slightly crenated, without node; vas deferens narrow and long; vagina cylindrical, long; oviduct shorter, proximally hardly tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria madurensis sp. nov. differs from the neighbouring Landouria intumescens in the presence of longish scales all over the teleoconch (Fig. 8), a wider umbilicus (U/D 0.38–0.40 vs 0.22–0.34 in L. intumescens) and a relatively long vagina (P:V 2.1 vs 2.4–3.9 in L. intumescens).

A second, more conical species of Landouria without longish scales and with a narrower umbilicus, of which only juveniles were found, lives sympatrically with Landouria madurensis sp. nov. in Nepa Monkey lowland rainforest on Madura Island.

Distribution

Landouria madurensis sp. nov. is endemic to Madura Island (Fig. 53).