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Published February 1, 2023 | Version v1
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Reproductive and Morphological Characteristics of Hemidactylus turcicus (Squamata: Gekkonidae) and Tarentola annularis (Squamata: Phyllodactylidae) in Southern Egypt

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Akef, Mamdouh S. (2023): Reproductive and Morphological Characteristics of Hemidactylus turcicus (Squamata: Gekkonidae) and Tarentola annularis (Squamata: Phyllodactylidae) in Southern Egypt. Ichthyology & Herpetology 111 (1): 29-43, DOI: 10.1643/h2021109, URL: http://dx.doi.org/10.1643/h2021109

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urn:lsid:plazi.org:pub:FF93FF8EFF964C15FF963442FFFCFFE2

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Figure: 10.5281/zenodo.7846998 (DOI)
Dataset: http://table.plazi.org/id/DF7C6668FF954C16FF8D3619FD74FD6B (URL)
Dataset: http://table.plazi.org/id/DF7C6668FF924C11FF1834F0FD01FF03 (URL)
Figure: 10.5281/zenodo.7847000 (DOI)
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Figure: 10.5281/zenodo.7847002 (DOI)
Figure: 10.5281/zenodo.7847004 (DOI)

References

  • Akef, M. S. 2022. Sexual size dimorphism and female reproductive and fat body cycles of the viviparous Chalcides ocellatus and Sphenops sepsoides (Squamata: Scincidae) from southern Egypt. Ichthyology and Herpetology 110:237-251.
  • Aldridge, R. D., D. S. Siegel, S. R. Goldberg, and R. A. Pyron. 2020. Seasonal timing of spermatogenesis and mating in squamates: a reinterpretation. Copeia 108:231- 264.
  • Angelini, F., G. Ciarcia, O. Picariello, and E. D'Alterio. 1983. The annual spermatogenetic cycle of Tarentola mauritanica L. (Reptilia, Gekkonidae). I-The spermatogenetic cycle in nature. Amphibia-Reptilia 4:171-184I
  • Annable, T. J. 2004. Reproductive biology of Nephrurus and Underwoodisaurus geckos (Reptilia: Gekkonidae: Carphodactylini). Unpubl. Ph.D. diss., University of Sydney, Sydney, Australia.
  • Aragon, P., P. Lopez, and J. Martin. 2006. Roles of male residence and relative size in the social behavior of Iberian rock lizards, Lacerta monticola. Behavioral Ecology and Sociobiology 59:762-769.
  • Athar, H. S., S. N. Hasnain, and M . Zain-Ul-Abedin. 1973. Studies on carbohydrate metabolism in lizards. Zeitschrift fur Naturforschung. Teil B. Anorganische Chemie, organische Chemie, Biochemie, Biophysik, Biologie 27:1531- 1535.
  • Atzori, A., F. Berti, T. Cencetti, S. Fornasiero, M. Tamburini, and A. L. Zuffi. 2007. Advances in methodologies of sexing and marking less dimorphic gekkonid lizards: the study case of the Moorish gecko, Tarentola mauritanica. Amphibia-Reptilia 28:449-454.
  • Barbault, R., and Y. P. Mou. 1988. Population dynamics of the common wall lizard, Podarcis muralis, in southwestern France. Herpetologica 44:38-47.
  • Birkhead, T. R., and A. P. MOller. 1993. Sexual selection and the temporal separation of reproductive events-sperm storage data from reptiles, birds and mammals. Biological Journal of the Linnean Society 50:295-311.
  • Bonneaud, C., E. Marnocha, A. Herrel, B. Vanhooydonck, D. J. Irschick, and T. B. Smith. 2016. Developmental plasticity affects sexual size dimorphism in an anole lizard. Functional Ecology 30:235-243.
  • Brandt, R., and C. A. Navas. 2011. Life-history evolution on Tropidurinae lizards: influence of lineage, body size and climate. PLoS ONE 6:e20040.
  • Bull, C. M., A. Main, and G. Dubas. 1991. Asynchronous seasonal activity of male and female sleepy lizards, Tiliqua rugosa. Journal of Herpetology 25:436-441.
  • Bustard, H. R. 1968. The egg-shell of gekkonid lizards: a taxonomic adjunct. Copeia 1968:162-164.
  • Church, G. 1962. The reproductive cycles of the Javanese house geckos, Cosymbotus platyurus, Hemidactylus frenatus, and Peropus mutilatus. Copeia 1962:262-269.
  • Cox, R. M., D. S. Stenquist, and R. Calsbeek. 2009. Testosterone, growth and the evolution of sexual size dimorphism. Journal of Evolutionary Biology 22:1586- 1598.
  • Crews, D., and M. C. Moore. 1986. Evolution of mechanisms controlling mating behavior. Science 231:121-125.
  • Crochet, P. A., and J. P. Renoult. 2008. Tarentola annularis annularis (Geoffroy de Saint-Hilaire, 1827) preying on a mammal. Herpetology Notes 1:58-59.
  • Curlis, J. D., C. L. Cox, and R. M. Cox. 2021. Sex-specific population differences in resting metabolism are associated with intraspecific variation in sexual size dimorphism of brown anoles. Physiological and Biochemical Zoology 94: 205-214.
  • Davis, J., and R. G. Ford. 1983. Home range in the western fence lizard (Sceloporus occidentalis occidentalis). Copeia 1983:933-940.
  • De Fuentes-Fernandez, M., M. M. Suarez-Rancel, and M. Molina-Borja. 2016. Variation in body size and morphometric traits of males and females of the wall gecko, Tarentola delalandii (Phyllodactylidae) from different environments on Tenerife. African Journal of Herpetology 65: 83-98.
  • Derickson, W. K. 1976. Lipid storage and utilization in reptiles. American Zoology 16:711-723.
  • Dial, B. E., and L. L. Grismer. 1992. A phylogenetic analysis of physiological-ecological character evolution in the lizard genus Coleonyx and its implications for historical biogeographic reconstruction. Systematic Biology 41:178-195.
  • Doligez, B., and J. Clobert. 2003. Clutch size reduction as a response to increased nest predation rate in the Collared Flycatcher. Ecology 84:2582-2588.
  • Eckstut, M. E., E. R. Lemons, and D. M. Sever. 2009. Annual dynamics of sperm production and storage in the Mediterranean gecko, Hemidactylus turcicus, in the southeastern United States. Amphibia-Reptilia 30:45-56.
  • Fitch, H. S. 1970. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication 52:1-247.
  • Ford, N., and R. Seigel. 1994. An experimental study of the trade-offs between age and size at maturity: effects of energy availability. Functional Ecology 8:91-96.
  • Fox, H. 1977. The urogenital system of reptiles, p. 1-157. In: Biology of the Reptilia. Volume 6. C. Gans and T. S. Parsons (eds.). Academic Press, London and New York.
  • Fox, R. J., L. Fromhage, and M. D. Jennions. 2019. Sexual selection, phenotypic plasticity and female reproductive output. Philosophical Transactions of the Royal Society B 374:20180184.
  • Galan, P. 1996. Sexual maturity in a population of the lacertid lizard Podarcis bocagei. Herpetological Journal 6:87- 93.
  • Goodwin, N. B., N. K. Dulvy, and J. D. Reynolds. 2002. Life-history correlates of the evolution of live bearing in fishes. Philosophical Transactions of the Royal Society B 357:259- 267.
  • Greer, A. E. 1989. The Biology and Evolution of Australian Lizards. Surrey Beatty, Chipping Norton, New South Wales.
  • Hernandez-Franyutti, A., and M. C. Uribe. 2012. Seasonal spermatogenic cycle and morphology of germ cells in the viviparous lizard Mabuya brachypoda (Squamata, Scincidae). Journal of Morphology 273:1199-1213.
  • Hernandez-Gallegos, O., F. R. Mendez-de la Cruz, M. Villagran-SantaCruz, J. L. Rheubert, G. Granados-Gonzalez, and K. M. Gribbins. 2015. Seasonal spermatogenesis in the Mexican endemic oviparous lizard, Sceloporus aeneus (Squamata: Phrynosomatidae). Spermatogenesis 4: e988585.
  • Herrel, A., L. Spithoven, R. van Damme, and F. de Vree. 1999. Sexual dimorphism of head size in Gallotia galloti: testing the niche divergence hypothesis by functional analyses. Functional Ecology 13:289-297.
  • Hibbitts, T. J., E. R. Pianka, R. B. Huey, and M. J. Whiting. 2005. Ecology of the common barking gecko (Ptenopus garrulus) in Southern Africa. Journal of Herpetology 39: 509-515.
  • Higham, T. E., P. Korchari, and L. D. McBrayer. 2011. How to climb a tree: lizards accelerate faster, but pause more, when escaping on vertical surfaces. Biological Journal of the Linnean Society 102:83-90.
  • Huang, W. S. 2006. Ecological characteristics of the skink, Mabuya longicaudata, on a tropical East Asian island. Copeia 2006:293-300.
  • Ibarguengoytia, N. R., and L. M. Casalins. 2007. Reproductive biology of the southernmost gecko Homonota darwini: convergent life-history patterns among Southern Hemisphere reptiles living in harsh environments. Journal of Herpetology 41:72-80.
  • Ibrahim, A. 2004. Behavioural ecology of the white-spotted gecko, Tarentola annularis (Reptilia: Gekkonidae), in Ismailia City, Egypt. Zoology in the Middle East 31:23-38.
  • Ikeuchi, I. 2004. Male and female reproductive cycles of the Japanese gecko, Gekko japonicus, in Kyoto, Japan. Journal of Herpetology 38:269-274.
  • James, C., and R. Shine. 1985. The seasonal timing of reproduction a tropical-temperate comparison in Australian lizards. Oecologia 67:464-474.
  • Johnson, J. B., L. D. McBrayer, and D. Saenz. 2005. Allometry, sexual size dimorphism, and niche partitioning in the Mediterranean gecko (Hemidactylus turcicus). The Southwestern Naturalist 50:435-439.
  • Johnston, G., and A. Bouskila. 2007. Sexual dimorphism and ecology of the gecko, Ptyodactylus guttatus. Journal of Herpetology 41:506-513.
  • Jones, S. M., and R. E. Ballinger. 1987. Comparative life histories of Holbrookia maculate and Sceloporus undulatus in western Nebraska. Ecology 68:1828-1838.
  • Kelada, M., C. Moulton, C. Nguyen, and G. R. Olague. 2021. Ventral coloration and body condition do not affect territorial behavior in two Sceloporus lizards. California Ecology and Conservation Research 5:1-8.
  • Kingsolver, J. G., and D. W. Pfennig. 2004. Individual-level selection as a cause of Cope's rule of phyletic size increase. Evolution 58:1608-1612.
  • Kratochvil, L., and D. Frynta. 2006. Egg shape and size allometry in geckos (Squamata: Gekkota), lizards with contrasting eggshell structure: why lay spherical eggs? Journal of Zoological Systematics and Evolutionary Research 44:217-222.
  • Kumar, S., B. Roy, and U. Rai. 2011. Hormonal regulation of testicular functions in reptiles, p. 63-88. In: Hormones and Reproduction of Vertebrates. Volume 3. D. O. Norris and K. H. Lopez (eds.). Elsevier, Academic Press, San Diego.
  • Liang, T., L. Shi, G. Bempah, and C. H. Lu. 2021. Sexual size dimorphism and its allometry in Chinese lizards. Evolutionary Ecology 35:323-335.
  • Licht, P., H. E. Hoyer, and P. G. W. J. Oordt. 2009. Influence of photoperiod and temperature on testicular recrudescence and body growth in the lizards Lacerta sicula and Lacerta muralis. Journal of Zoology 157:469-501.
  • Lopez-Alcaide, S., L. Y. Cuateta-Bonilla, and R. Macip- Rios. 2020. Reproductive output in spiny lizards (genus Sceloporus) with different reproductive mode. A comparative approach. Revista Mexicana de Biodiversidad 91: e913020.
  • Losos, J. B. 1990. The evolution of form and function: morphology and locomotor performance in West Indian Anolis lizards. Evolution 44:1189-1203.
  • Marion, K. R. 1982. Reproductive cues for gonadal development in temperate reptiles: temperature and photoperiod effects on the testicular cycle of the lizard Sceloporus undulates. Herpetologica 38:26-39.
  • Mathies, T., J. A. Cruz, V. A. Lance, and J. A. Savidge. 2010. Reproductive biology of male brown tree snakes (Boiga irregularis) on Guam. Journal of Herpetology 44:209-221.
  • Meiri, S. 2011. Bergmann's rule-what's in a name? Global Ecology and Biogeography 20:203-207.
  • Meshaka, W. E., Jr. 1995. Reproductive cycle and colonization ability of the Mediterranean gecko (Hemidactylus turcicus) in south-central Florida. Florida Scientist 58:10- 15.
  • Mesquita, D., G. C. Costa, R. Colli, T. B. Costa, D. B. Shepard, L. J. Vitt, and E. R. Pianka. 2016. Life-history patterns of lizards of the world. The American Naturalist 187:689-705.
  • Meylan, P. A., R. Schuler, P. Moler, and R. M. Wood. 2002. Spermatogenic cycle of the Florida softshell turtle, Apalone ferox. Copeia 2002:779-786.
  • Nogueira, K. O. P. C., S. S. R. Sartori, V. A. Araujo, C. A. Neves, and B. Kolisnyk. 2015. Sperm storage in Hemidactylus mabouia: morphological and ultrastructural aspects of a reproductive. Animal Reproductive Science 159: 212-216.
  • Olsson, M., R. Shine, E. Wapstra, B. Ujvari, and T. Madsen. 2002. Sexual dimorphism in lizard body shape: the roles of sexual selection and fecundity selection. Evolution 56: 1538-1542.
  • Packard, M. J., G. C. Packard, and T. J. Boardman. 1982. Structure of eggshells and water relations of reptilian eggs. Herpetologica 38:136-155.
  • Pafilis, P., S. Meiri, J. Foufopoulos, and E. Valakos. 2009. Intraspecific competition and high food availability are associated with insular gigantism in a lizard. Naturwissenschaften 96:1107-1113.
  • Parker, G. A. 1982. Why are there so many tiny sperm? Sperm competition and the maintenance of two sexes. Journal of Theoretical Biology 96:281-294.
  • Paulissen, M. A., and T. M. Buchanan. 1991. Observations on the natural history of the Mediterranean gecko, Hemidactylus turcicus (Sauria: Gekkonidae) in northwest Arkansas. Journal of the Arkansas Academy of Science 45:81-83.
  • Paz, M. M., N. E. Garcia, R. V. Semhan, F. J. Lobo, and C. S. Abdala. 2019. Study of lipid reserves in Liolaemus koslowskyi (Squamata: Liolaemidae): reproductive and ecological implications. Journal of Comparative Physiology B 189:595-609.
  • Perry, G. 1996. The evolution of sexual dimorphism in the lizards Anolis polylepis (Iguania): evidence from intraspecific variation in foraging behavior and diet. Canadian Journal of Zoology 74:1238-1245.
  • Pincheira-Donoso, D., and J. Hunt. 2017. Fecundity selection theory: concepts and evidence. Biological Reviews 92:341-356.
  • Prasad, M. R. N., and M. K. Sanyal. 1969. Effect of sex hormone on the sexual segment of the kidney and other accessory reproductive organs of the Indian house lizard Hemidactylus flaviviridis Ruppell. General and Comparative Endocrinology 12:110-118.
  • Punzo, F. 2001. The Mediterranean gecko, Hemidactylus turcicus: life in an urban landscape. Florida Scientist 64: 56-66.
  • Ramirez-Bautista, A., D. Hernandez-Ramos, A. R. Martinez, and J. C. Marshall. 2009. Fat bodies and liver mass cycles in Sceloporus grammicus (Squamata: Phrynosomatidae) from southern Hidalgo, Mexico. Herpetological Conservation and Biology 4:164-170.
  • Reddy, P. M., V. R. Reddy, C. V. Reddy, and S. P. Rap. 1979. Egg weight, shape index and hatchability in khaki Campbell duck egg. Indian Journal of Poultry Science 14:26-31.
  • Rheubert, J. L., C. M. Murray, D. S. Siegel, J. Babin, and D. M. Sever. 2011. The sexual segment of Hemidactylus turcicus and the evolution of sexual segment location in squamata. Journal of Morphology 272:802-813.
  • Rheubert, J. L., E. H. Poldemann, M. E. Eckstut, M. H. Collier, D. M. Sever, and K. M. Gribbins. 2009. Temporal germ cell development strategy during mixed spermatogenesis within the male Mediterranean gecko, Hemidactylus turcicus (Reptilia: Gekkonidae). Copeia 2009:793-800.
  • Rheubert, J. L., K. T. Touzinsky, D. M. Sever, R. D. Aldridge, A. J. Wilmes, D. S. Siegel, and K. M. Gribbins. 2014. Reproductive biology of Sceloporus consobrinus (Phrynosomatidae): male germ cell development and reproductive cycle comparisons within spiny lizards. Journal of Herpetology 48:162-171.
  • Righi, A. F., L. B. Nascimento, and C. A. B. Galdino. 2012. Seasonal reproduction in the rock gecko Phyllopezus pollicaris from a rock field habitat in southeastern Brazil. Journal of Herpetology 46:632-636.
  • Rocha, C. F. D. 1992. Reproductive and fat body cycles of the tropical sand lizard (Liolaemus lutzae) of southeastern Brazil. Journal of Herpetology 26:17-32.
  • Roitberg, E. S., V. F. Orlova, N. A. Bulakhova, V. N. Kuranova, G. V. Eplanova, O. I. Zinenko, O. Arribas, L. Kratochvil, K. Ljubisavljevic, V. P. Starikov, H. Strijbosch, S. Hofmann, O. A. Leontyeva, and W. Bohme. 2020. Variation in body size and sexual size dimorphism in the most widely ranging lizard: testing the effects of reproductive mode and climate. Ecology and Evolution 10: 4531-4561.
  • Rose, F. L., and C. D. Barbour. 1968. Ecology and reproductive cycles of the introduced gecko, Hemidactylus turcicus, in the southern United States. The American Midland Naturalist 79:159-168.
  • Runemark, A., K. Sagonas, and E. Svensson. 2015. Ecological explanations to island gigantism: dietary niche divergence, predation, and size in an endemic lizard. Ecology 96:2077-2092.
  • Saenz, D. 1996. Dietary overview of Hemidactylus turcicus with possible implications of food partitioning. Journal of Herpetology 30:461-466.
  • Sanyal, M. K., and M. R. N. Prasad. 1967. Reproductive cycle of the Indian house lizard, Hemidactylus flaviviridis Ruppell. Copeia 1967:627-633.
  • Schluter, D. 2001. Ecology and the origin of species. Trends in Ecology and Evolution 16:372-380.
  • Selcer, K. W. 1986. Life history of a successful colonizer: the Mediterranean Gecko, Hemidactylus turcicus, in southern Texas. Copeia 1986:956-962.
  • Selcer, K. W. 1987. Seasonal variation in fat body and liver mass on the introduced Mediterranean gecko, Hemidactylus turcicus. Journal of Herpetology 21:74-78.
  • Selcer, K. W. 1990. Egg-size relationships in a lizard with fixed clutch size: variation in a population of the Mediterranean gecko. Herpetologica 46:15-21.
  • Selcer, K. W. 1992. Lipid storage during formation of early- and late-season clutches in Hemidactylus turcicus. Journal of Herpetology 26:209-213.
  • Sever, D. M., and W. A. Hopkins. 2004. Oviductal sperm storage in the ground skink Scincella laterale Holbrook (Reptilia: Scincidae). Journal of Experimental Zoology 301A:599-611.
  • Shine, R. 1988. The evolution of large body size in females: a critique of Darwin's ''fecundity advantage'' model. The American Naturalist 131:124-131.
  • Shine, R. 1992. Relative clutch mass and body shape in lizards and snakes: is reproductive investment constrained or optimized? Evolution 46:828-833.
  • Siegel, D. S., A. Miralles, J. L. Rheubert, and D. M. Sever. 2015. Female reproductive anatomy: cloaca, oviduct and sperm storage, p. 144-195. In: Reproductive Biology and Phylogeny of Lizards and Tuatara. J. L. Rheubert, D. S. Siegel, and S. E. Trauth (eds.). CRC Press, Boca Raton, Florida.
  • Simbotwe, M. P. 1980. Reproductive biology of the skinks Mabuya striata and Mabuya quinquetaeniata in Zambia. Herpetologica 36:99-104.
  • Sion, G., M. J. Watson, and A. Bouskila. 2021. Measuring body condition of lizards: a comparison between noninvasive dual energy X-ray absorptiometry, chemical fat extraction and calculated indices. Frontiers in Zoology 18: 1.
  • Sparkman, A. M., A. D. Clark, L. J. Brummett, K. R. Chism, L. L. Combrink, N. M. Kabey, and T. S. Schwartz. 2018. Convergence in reduced body size, head size, and blood glucose in three island reptiles. Ecology and Evolution 8: 6169-6182.
  • Stabler, L. B., W. L. Johnson, K. J. Locey, and P. A. Stone. 2012. A comparison of Mediterranean Gecko (Hemidactylus turcicus) populations in two temperate zone urban habitats. Urban Ecosystem 15:653-666.
  • Stamps, J. A., M. Mangel, and J. A. Phillips. 1998. A new look at relationships between size at maturity and asymptotic size. The American Naturalist 152:470-479.
  • Starostova, Z., L. Kubica, and L. Kratochvil. 2010. Macroevolutionary pattern of sexual size dimorphism in geckos corresponds to intraspecifc temperature-induced variation. Journal of Evolutionary Biology 23:670-677.
  • Uller, T., T. Schwartz, T. Koglin, and M. Olsson. 2013. Sperm storage and sperm competition across ovarian cycles in the dragon lizard, Ctenophorus fordi. Journal of Experimental Zoology 319A:404-408.
  • Vanhooydonck, B., A. Andronescu, A. Herrel, and D. J. Irschick. 2005. Effects of substrate structure on speed and acceleration capacity in climbing geckos. Biological Journal of the Linnean Society 85:385-393.
  • Velasco, J. A., F. Villalobos, J. A. F. Diniz-Filho, S. Poe, and O. Flores-Villela. 2020. Macroecology and macroevolution of body size in Anolis lizards. Ecography 43:812-822.
  • Veldsman, L. M., H. Kylin, P. Bronkhorst, I. Engelbrecht, and H. Bouwman. 2020. A method to determine the
  • Vitt, L. J. 1986. Reproductive tactics of sympatric gekkonid lizards with a comment on the evolutionary and ecological consequences of invariant clutch size. Copeia 1986:773- 786.
  • Vitt, L. J., and H. J. Price. 1982. Ecological and evolutionary determinants of relative clutch mass in lizards. Herpetologica 38:237-255.
  • Warner, D. A., M. S. Johnson, and T. R. Nagy. 2016. Validation of body condition Indices and quantitative magnetic resonance in estimating body composition in a small lizard. Journal of Experimental Zoology Part A: Ecological Genetics and Physiology 325:588-597.
  • Warren, D. C., and R. L. Schnepel. 1940. The effect of air temperatures on egg shell thickness in the fowl. Poultry Science 19:67-72.
  • Wegener, J. E., K. P. Mulder, R. M. Pringle, J. B. Losos, and J. J. Kolbe. 2019. Head size of male and female lizards increases with population density across island populations in the Bahamas. Breviora 566:1-9.
  • Werner, Y. L. 1989. Egg size and egg shape in near-eastern gekkonid lizards. Israel Journal of Zoology 35:199-213.
  • Whittier, J. M., and R. R. Tokarz. 1992. Physiological regulation of sexual behavior in female reptiles, p. 24-69. In: Hormones, Brain, and Behavior: Biology of the Reptilia. Volume 18. C. D. Gans and D. Crews (eds.). University of Chicago Press, Chicago.
  • Wormington, J. D., and S. A. Juliano. 2014. Sexually dimorphic body size and development time plasticity in Aedes mosquitoes (Diptera: Culicidae). Evolutionary Ecology Research 16:223-234.
  • Zaworski, J. P. 1992. Reproduction in the gecko genus Tarentola. Palt 1. Captive breeding and husbandry of the Egyptian white spotted gecko, Tarentola annularis (Geoffroy). Dactylus 1:24-31.
  • Zhang, Y.-P., W.-G. Du, and L.-J. Zhu. 2009. Differences in body size and female reproductive traits between two sympatric geckos, Gekko japonicus and Gekko hokouensis. Folia Zoology 58:113-122.