Lanonia poilanei Henderson & Nguy

17. Lanonia poilanei Henderson & Nguy ễn Quốc Dựng (2017: 161). Type:— VIETNAM. Khanh Hoa: new road from Nha Trang to Da Lat, 12.232N 108.775E, ca. 550 m, 9 July 2014, A. Henderson & Nguyễn Qu ốc Dựng 3936 (holotype VFM!, isotype NY!).

Stems 1.3(0.8–2.0) m long, 3.8(3.5–4.0) cm diameter, solitary or clustered. Leaves number not recorded; ligules 9.0 cm long, soon disintegrating into fibers; sheaths and proximalmost part of petioles with scattered, brown scales; petioles 95.0(80.0–110.0) cm long, 0.6(0.6–0.7) cm wide at the apex; petiole thorns usually poorly developed, brown or black, more or less regularly arranged on proximalmost part of petiole; hastulas flat, rounded, not infolded; leaf blades 55.0 cm wide; costas 12.3(9.7–13.8) cm long, narrow, with a pulvinus at the apex abaxially, with the numerous segments free to the base except the middle pair joined at their bases; segments 20(19–21) per leaf, not mottled, with minute, reddish-brown scales abaxially, with (on dried specimens) straight margins; middle segments 48.3(33.5–57.5) cm long, 6.3(4.5–7.5) cm wide at the apex; apices of middle segments with adaxial splits not much deeper than abaxial ones. Inflorescences 70.0 cm long; prophylls and peduncles length not recorded; rachis bracts flattened, splitting apically and laterally, almost woody and scarcely tomentose; rachillae not filiform nor thick and ribbed, sparsely to moderately covered with short, mostly simple, brownish hairs; staminate inflorescences not recorded; pistillate rachises absent or 18.0 cm long; pistillate partial inflorescences 1-2, branched to 3 orders; pistillate rachillae on proximalmost partial inflorescence 11(9–15), 11.9(9.2–13.5) cm long, 1.6(1.3–1.8) mm diameter; pistillate flowers length not recorded, with short styles; fruits 8.7 mm long, 6.4 mm diameter, globose, red, with smooth surfaces; seeds with a curved basal intrusion.

Distribution and habitat:—Southern Vietnam in Khanh Hoa province in lowland rainforest at 576(400–800) m elevation (Fig. 17).

Taxonomic notes:— Specimens determined as preliminary species Lanonia poilanei were polymorphic for stem branching, hastula shape, and segment splitting. Therefore the preliminary species was split into three species such that variables were consistent within each one (except that stem branching was treated as a trait), and these three are recognized as phylogenetic species, L. poilanei, L. honheoensis, and L. nuichuaensis.

Two specimens (Henderson 3695, 3696) from Hon Ba were included in Lanonia poilanei by Henderson & Nguyễn Quốc Dựng (2017), as well as two other similar specimens (Chevalier 38844, 38730) from the same locality. All are sterile and remain unidentified. Two other sterile specimens (Henderson 4067, 4107), somewhat similar to those of L. poilanei, also remain unidentified.