Anocha japonica Jaschhof, 2017, sp. nov.

Anocha japonica sp. nov.

Fig. 3A–B

Diagnosis. The eye-bridge is incomplete; veins posterior to R4+5 are so weak that they are indiscernible in the only specimen available of this species. Male genitalic structures typical of Anocha japonica are the subcylindrical, almost straight gonostyli provided with a long, pale apical tooth, and the gonocoxites, which are as long as wide and have a deeply V-shaped ventral emargination (Fig. 3A). Females are unknown.

Differential diagnosis. Anocha japonica differs from A. minuta, a species with similar male genitalia (Jaschhof & Jaschhof 2009: fig. 21C), as follows: the scape is setose (asetose in A. minuta); the scutum has both lateral and dorsocentral setae (only lateral setae); the wing membrane is fully setose (almost asetose); the posterior wing veins are indistinct (distinct); and the toothed claws are single-pointed apically (toothless, two-pointed).

Other male characters. Body size 2.1 mm. Head. Eye-bridge 5 ommatidia long at vertex. Postocular bristles absent. Flagellomeral nodes with sparse microtrichia on basal halves (not drawn in Fig. 3B). Neck of fourth flagellomere 1.8 times longer than node (Fig. 3B). Apical palpal segment subcylindrical, not club-shaped. Terminalia (Fig. 3A). Ninth tergite small, posterior edge sinuous (apparently no artifact of preparation). Gonocoxites: membranous window narrow, ventrobasal edge slightly convex. Gonostylus twice as long as wide, pointing dorsomedially; apical claw protruding, not adpressed. Tegmen subtrapezoid. Aedeagal teeth about 10, slightly larger than in other Anocha.

Etymology. The species epithet, japonica, is an adjective referring to Japan, the only distribution known of this species.

Holotype. Male (no. CEC403), Japan, Honshu, Ibaraki, Kitaibaraki, Sadanami, Ogawa Research Forest, mature broadleaf forest, 1 Dec. 1999 – 2 March 2000, Malaise trap, M. & C. Jaschhof (in KUEC).

Distribution and phenology. Japan (Honshu). The only specimen known of Anocha japonica was collected in winter in a mature broadleaf deciduous forest.