Noideattella lakana Álvarez-Padilla & Ubick & Griswold 2012, new species

Noideattella lakana, new species Figures 257–284, 375

TYPES: Male holotype and female allotype from Berlese and sifted leaf litter of a tropical dry forest at an elevation of 325 m at Reserve Speciale d’Ambre, 3.5 km 235° SW Sakaramy, 12°28′8″S, 49°14′32″E, Antsiranana Province, 26–31 January 2001, Fisher-Griswold Arthropod Team, deposited in CASC (PBI_OON 00003275). The species epithet, a noun in apposition, is taken from the Malagasy word for a dugout canoe or any small boat .

DIAGNOSIS: N. lakana can be differentiated from other Noideattella species by the following combination of characters: habitus color dark red-brown, body length 2.1 to 2.4 mm; carapace surface of elevated portion of pars cephalica strongly granulate, sides granulate (figs. 260, 275, 276); sternum furrows with row of large pits, surface coarsely reticulate, with microsculpture medially and in furrows (fig. 277); book lung covers large and elliptical (figs. 258, 259).

DESCRIPTION: Male (PBI_OON 0003462): Total length 2.1. Cephalothorax: Carapace dark red-brown, sternum longer than wide, dark red-brown, anterior margin with continuous transverse groove (fig. 273). Mouthparts: Chelicerae, endites, and labium dark red-brown, paturon inner margin with scattered setae (fig. 263). Abdomen: Ovoid. Book lung covers large, elliptical. Dorsal scutum dark red-brown, middle surface smooth, sides smooth. Postepigastric scutum orange-brown, long, semicircular. Male genital opening slightly protruded, epigastric scutum apodemes small, separated from the anterior book lung opening and connected by a furrow. Posterior spiracles connected by a groove (fig. 268). Legs: Orange-brown, without color pattern. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I, II p3-1-1; r2-1-1; IV p0-0-1; metatarsi: I, II p1-1-0; r1-1-0. Tarsal organ with two sensilla visible. Genitalia: Epigastric region with sperm pore situated at level of anterior spiracles (fig. 268). Palp proximal segments pale orange; cymbium pale orange, with distal patch of setae; bulb pale orange (figs. 262, 264). Embolus dorsal and ventral lamellae with at least three distal apophyses each (figs. 281–284). Palp as in figures 279 and 280. Female (PBI_ OON 0003462): Total length 2.4. As in male except as noted. Legs: Metatarsi III and IV weak ventral scopula absent. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I, II p3-1-1; r3-1-1; IV p0-0-1; metatarsi: I, II p1-1-0; r1-1-0. Trichobothria of metatarsus: I and IV one. Genitalia: Genital opening slit shaped (fig. 267). Sperm sac secretory glands openings flat, extending all over the receptaculum posterior surface. GAp process shaped as a twisted tube (figs. 269–271). Genitalia as in figures 265 and 266.

MATERIAL EXAMINED: N = 86. Madagascar: Antananarivo: 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, montane rainforest, 1300 m, S18.47333°, E47.96000°, Dec. 05–13, 2000, Fisher-Griswold Arthro- pod Team, 1♀ (CASC PBI _ OON 3941). Antsiranana: 7.2 km 142° SE Antsiranana (= Diego Suarez), Montagne des Francais, 180 m, S12.53796°, E49.56361°, Feb. 23–25, 2001, L.J. Boutin, 1♀ (CASC PBI _ OON 3921); Forêt de Binara, 9.4 km 235° SW Daraina, montane rainforest, 1100 m, S13.26333°, E49.60000°, Dec. 05, 2003, B.L. Fisher, 5 mixed (CASC PBI _ OON 3153); P.N. Montagne d’Ambre, 2.79 air km NE of park entrance forest, 1000 m, S12.53333°, E49.16666°, Nov. 21–30, 1993, J. Coddington, S. Larcher, C. Griswold, R. Andriamasimanana, N. Scharff, 1♀ (CASC PBI _ OON 3460); 12 mixed (ZMUC PBI _ OON 3462); 5 mixed (CASC PBI _ OON 3463); 5 mixed (CASC PBI _ OON 3465); 2♂ (CASC PBI _ OON 3468); 2♀ (CASC PBI _ OON 3473); 1♂ (CASC PBI _ OON 3474); 1♂ (CASC PBI _ OON 3475); 15 mixed (CASC PBI _ OON 3477); 2 mixed (CASC PBI _ OON 36378); 5 mixed (CASC PBI _ OON 36380); 13 mixed (CASC PBI _ OON 36389); 6 mixed (CASC PBI _ OON 36392); Parc National Montagne d’Ambre, 3.6 km 235° SW Joffreville, montane rainforest, 925 m, S12.53444°, E49.17944°, Jan. 20–26, 2001, Fisher-Griswold Arthropod Team, 4 mixed (CASC PBI _ OON 36376); Reserve Speciale d’Ambre, 3.5 km 235° SW Sakaramy, tropical dry forest, 325 m, S12.46888°, E49.24222°, Jan. 26–31, 2001, Fisher-Griswold Arthropod Team, 1♀ (CASC PBI _ OON 3359); Reserve Speciale de l’Ankarana, 22.9 km 224° SW Anivorano Nord, tropical dry forest, 80 m, S12.90888°, E49.10972°, Feb. 10–16, 2001, Fisher-Griswold Arthropod Team, 1♂ (CASC PBI _ OON 3172); Reserve Speciale l’Ankarana, 13.6 km 192° SSW Anivorano Nord, tropical dry forest, 210 m, S12.86361°, E49.22583°, Feb. 16–20, 2001, Fisher-Griswold Arthropod Team, 3 mixed (CASC PBI _ OON 3721).