Speocera ankalana Li & Li 2019, sp. nov.

Speocera ankalana Li & Li sp. nov. (Figs 3–4, 55B)

Material examined. Holotype ♂ (IZCAS), Madagascar, Antsiranana Province, Ambilobe District, Ankalana National Park (12º57.010′S, 49º07.021′E), elev. 140 m, 16.XII.2016, leg. H. Zhao. Paratype 1♀ (IZCAS), same data as holotype.

Etymology. The specific name is derived from the type locality; noun.

Diagnosis. The new species is similar to S. transleuser Deeleman-Reinhold, 1995, but males can be distinguished by the laterally strongly excavated chelicerae (Fig. 55B) and the elliptical palpal bulb (Figs 3A–B), and the females by the very short, straight afferent spermathecae (Fig. 4A). S. transleuser males have antero-posteriorly compressed chelicerae and a nearly flattened palpal bulb, and females have long, curved afferent spermathecae (Deeleman-Reinhold, 1995: figs 257– 260).

Description. Male. Total length 0.96; carapace length 0.47; carapace width 0.41; abdomen length 0.45; abdomen width 0.28. Prosoma: carapace pyriform, narrowing gradually anteriorly, anterior margin smoothly curved in dorsal view; light yellow, suffused with purplish pattern (Fig. 4C); pars cephalica flat; fovea not visible. Sternum light yellow, suffused with purplish color, as long as wide. Endites light yellow suffused with purplish tinge; with serrula; labium light yellow suffused with purplish tinge. Clypeus nearly vertical; short. Chelicerae light yellow; external margin strongly excavated; promargin with a series of six teeth reaching the lamina; retromargin with two very small denticles (Fig. 55B). Eyes: six contiguous eyes surrounded by black pigmentation, PME smallest, PLE largest (Fig. 4C). Opisthosoma oval; purplish. Legs missing. Genitalia (Fig. 3). Palpal femur long; palpal tibia elongated, about 1.6 times as long as cymbium (Figs 3A–B). Cymbium conical, without prolateral extension. Bulb elliptical, much longer than wide. Embolus short, black, associated sclerites beltlike; sperm duct initially medium sized, becoming very narrow distally (Fig. 3D).

Female. Total length 1.23; carapace length 0.48; carapace width 0.40; abdomen length 0.70; abdomen width 0.56. Similar to male in color and general features (Figs 4C–D) but larger. Chelicerae unmodified. Legs light yellow; I, II, IV missing; III 1.46 (0.41, 0.14, 0.38, 0.29, 0.23). Genitalia. Internal genitalia with afferent spermathecae short, straight, not extending past the sides of the abdomen; tip of the afferent spermathecae question mark shaped (Fig. 4B); spermathecae oval (Fig. 4A).

Distribution. Known only from the type locality.