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Published March 20, 2021 | Version v1
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Corrections and Additions to the Checklist of Bark and Ambrosia Beetles (Coleoptera: Curculionidae: Scolytinae, Platypodinae) of Mesic Montane Forest in Xalapa, Veracruz, Mexico

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Atkinson, Thomas H., Ibarra Juárez, Luís Arturo (2021): Corrections and Additions to the Checklist of Bark and Ambrosia Beetles (Coleoptera: Curculionidae: Scolytinae, Platypodinae) of Mesic Montane Forest in Xalapa, Veracruz, Mexico. The Coleopterists Bulletin 75 (1): 227-239, DOI: 10.1649/0010-065X-75.1.227, URL: http://dx.doi.org/10.1649/0010-065x-75.1.227

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urn:lsid:plazi.org:pub:0D28FF870D4D3B0CFFE4FF951352D831
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http://publication.plazi.org/id/0D28FF870D4D3B0CFFE4FF951352D831

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Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFF4BF9ED11B9D8C3 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FF6D10DED9A7 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FE0E17C1DA8C (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FD5617C7DB4D (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFC89FC0710B7DBE3 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FC421773DC0F (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FBD61044DCF2 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FB5D1025DD1E (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FAA010CFDD85 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0AFD21FA2F10B9DE11 (URL)
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Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4B3B0BFD21F9071215D920 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FE821146D9B0 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FE13125AD9DA (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FE64124FDA41 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FDE01160DBB8 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FC1A113EDC49 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FBEB123EDDB3 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FA1D125ADDDD (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21FA671155DE66 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFF21F9C91155DEF0 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFD3FFF571074D9D1 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFD3FFE731085DB65 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFD3FFCCF175CDB8E (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFD3FFC511085DCF9 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFD3FFB5B1042DDE3 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFD3FFA421050DE43 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D4A3B0BFD3FF9ED1744DEF3 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFF57126AD947 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFEE9123EDA7A (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFDC4123DDAE1 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFD4312ADDB0A (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFCD51251DB94 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFC3F1177DBC4 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFC6F12D9DC6E (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFBF11111DD02 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFAAC1245DDAF (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BFA3613AEDE39 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BF99813A0DEBC (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FF4BF9071013D917 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FEB9173CD99A (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FE2410CBDAB1 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FD1010D0DADB (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FD6510EADB65 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FCCF175ADB8F (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FC5110B0DD29 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FA8817C7DD56 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B04FD21FAF81080DE36 (URL)
Taxonomic treatment: http://treatment.plazi.org/id/F11187FF0D453B07FD21F9991733DDDE (URL)
Figure: 10.5281/zenodo.5335104 (DOI)
Figure: 10.5281/zenodo.5335108 (DOI)

References

  • 62. Araptus confinis (Blandford, 1904). Phloem feeder, no reported hosts. Humid lowland and intermediate elevations from eastern Mexico to Panama.
  • 63. Araptus deyrollei (Blandford, 1904). Pith borer, oligophagous? Collected in dried fruits of milkweed vine, related species also breed in milkweeds (Apocynaceae). Lowland and intermediate elevations from eastern Mexico to Honduras.
  • 64. Araptus tabogae (Blackman, 1942). Phloem feeder-pith borer, polyphagous? Known from unrelated hosts, all vines. Intermediate and lowland habitats (dry and humid) from central Mexico (both coasts) to Colombia.
  • 65. *Cryptocarenus seriatus Eggers, 1933. Pith borer, polyphagous. Known from a wide variety of hosts. Lowland habitats from Florida and southern Texas, both coasts of Mexico to Brazil and Peru. This species was listed in Cryphalini by Noguera M. and Atkinson (1990).
  • 66. Pityophthorus alni Blackman, 1942. Phloem feeder, monophagous. Specialist on Alnus (Betulaceae). Intermediate elevations in central Mexico.
  • 67. Pityophthorus atomus Wood, 1964. Phloem feeder, oligophagous. Breeds in composite shrubs. Intermediate elevations in central Mexico.
  • 68. Pityophthorus hylocuroides Wood, 1964. Phloem feeder, polyphagous? Known only from Hidalgo and Veracruz at intermediate elevations.
  • 69. Pityophthorus paulus Wood, 1964. Phloem feeder, monophagous. Breeds in shrubs of the genus Senecio (Asteraceae). Intermediate elevations in northern and central Mexico.
  • 70. Pityophthorus tutulus Bright, 1986. Phloem feeder, monophagous? Probably specialist on poison ivy (Anacardiaceae: Toxicodendron) as are related species. Known only from the type locality near Xalapa.
  • 71. Pseudopityophthorus granulifer Wood, 1967. Phloem feeder, monophagous. Specialist on oaks (Quercus). Intermediate elevations from central Mexico to Honduras.
  • 72. Pseudopityophthorus opacicollis Blackman, 1931. Phloem feeder, oligophagous. Like most species in the genus, oaks are the primary hosts. However, on several occasions successful breeding has been observed in alders (Alnus) and black cherry (Rosaceae: Prunus) in forests where oaks are present from southern Arizona to Chiapas.
  • 73. Pseudopityophthorus singularis Wood, 1971. Phloem feeder, monophagous. Specialist on oaks (Quercus). Forests at intermediate elevations where oaks are present from Durango and Nuevo Leon to Veracruz.
  • 74. Pseudopityophthorus virilis Wood, 1971. Phloem feeder, monophagous. Specialist on oaks (Quercus). Forests at intermediate elevations where oaks are present in Hidalgo and Veracruz.
  • 75. Pseudopityophthorus xalapae Wood, 1987. Phloem feeder, monophagous. Specialist on oaks (Quercus). Known only from the immediate region of Xalapa.
  • 76. Sphenoceros aztecus Wood, 1971. Phloem feeder, oligophagous. Breeds in various genera of Lauraceae. Reported as Araptus aztecus (Wood). Bright (2019) removed the genus Sphenoceros from synonymy with Araptus.
  • 77. Stegomerus mexicanus Wood, 1967. Phloem feeder, polyphagous? All collections from hosts have been from vines. Intermediate and lowland habitats (dry and humid) from Michoacan to Tabasco. This species was listed in Cryphalini by Noguera M. and Atkinson (1990).
  • 78. Dendrocranulus macilentus (Blandford, 1898). Herbaceous hosts, oligophagous? No reported hosts but all other species in this genus are specialists breeding in stems of native and cultivated Cucurbitaceae vines. Veracruz to Costa Rica.
  • 79. Dendrocranulus mexicanus Wood, 1986. Herbaceous hosts, oligophagous.Specialist breeding in stems of native and cultivated Cucurbitaceae vines. Known only from the type locality near Xalapa.
  • 80. Scolytodes maurus (Blandford, 1897). Pith borer, monophagous: pith of petioles of fallen leaves of various species of Cecropia (Urticaceae). Intermediate and lower elevations in humid forests from eastern Mexico to Venezuela and Ecuador.
  • 81. Hylesinus caseariae Wood, 1986. Phloem feeder, monophagous? Type series reported from Casearia (Salicaceae) but all other New World species in genus breed in ashes (Oleaceae: Fraxinus). Known only from the type locality near Xalapa.
  • 82. Xylechinus marmoratus Blandford, 1897. Phloem feeder, monophagous. Specialist in Oreopanax (Araliaceae). Mesic forest at intermediate elevations from Honduras, Guatemala and the eastern-facing slopes of the Mexican states of Oaxaca, Puebla and Veracruz.
  • 83. Chaetophloeus struthanthi Wood, 1967. Phloem feeder, oligophagous. Specialist on mistletoes (Santalaceae, Loranthaceae). Intermediate elevations in central Mexico.
  • Micracidini 84. Hylocurus atkinsoni Wood, 1987. Sapwood borer, oligophagous. Known hosts are legumes,
  • 85. Micracis festiva Wood, 1969. Sapwood borer, polyphagous. Known from a variety of hosts. Intermediate elevations from central Mexico to Honduras.
  • 86. Micracis lignator Blackman, 1928. Sapwood borer, monophagous. Specialist on oaks (Quercus). Pine-oak communities in the Sierra Madre Occidental (including southeastern Arizona), the Sierra Madre Oriental, and the Neovolcanic axis of central Mexico.
  • 87. Micracis ovata Wood, 1956. Sapwood borer, polyphagous? No reported hosts but most species in this genus are generalists. Intermediate elevations in the Sierra Madre Oriental of Mexico from Tamaulipas to Veracruz.
  • 88. *Micracisella opacithorax (Schedl, 1940). Pith borer, oligophagous? All known hosts are legumes. Low to intermediate elevations along coast of Gulf of Mexico from southern Texas to Veracruz.
  • 89. Phloeocleptus caudatus Wood, 1956. Phloem feeder, monophagous. All collections from known hosts are from Persea. Humid forests at intermediate elevations in eastern Puebla and Veracruz.
  • 90. Pseudothysanoes contrarius Wood, 1974. Phloem feeder, oligophagous? All known hosts are legumes. Humid forests at intermediate elevations from Veracruz to Chiapas.
  • 91. Pseudothysanoes fimbriatus Wood, 1982. Phloem feeder, monophagous. Specialist on mistletoes (Santalaceae: Phoradendron). Humid forests at intermediate elevations in eastern Puebla and Veracruz.
  • 92. Thysanoes fimbricornis LeConte, 1876. Sapwood borer, polyphagous. Reported from many unrelated hosts, but with a strong apparent bias for willows (Salicaceae: Salix). Southeastern United States to southern Mexico. In Mexico more common at intermediate elevations.
  • 93. Chramesus chapuisii LeConte, 1876. Phloem feeder, oligophagous. All US records are from Celtis (Cannabaceae). Disjunct distribution.Largest population found throughout the southeastern United States to Texas. A separate area is in the Sierra Madre Oriental of Mexico from Tamaulipas to Veracruz.
  • 94. Chramesus vastus Wood, 1967. Phloem feeder, monophagous? All reported hosts are species of Inga (Leguminosae). This species was reported as Chramesus rotundatus (Chapuis) based on preliminary identifications by Wood. Subsequent comparisons by the senior author confirm that this is C. vastus. Humid forests at intermediate elevations from Veracruz to Panama.
  • 95. Chramesus xalapae Atkinson, 1989. Phloem feeder, monophagous? Host is unknown, but related species breed in monocots. Known only from the type locality.
  • 96. Phloeotribus demessus Blandford, 1897. Phloem feeder, oligophagous. Generally found in Ficus (Moraceae), but has also been found in Croton. Intermediate and lower elevations in humid and seasonally dry forests from Tamaulipas and Jalisco to Panama.
  • 97. Phloeotribus pruni Wood, 1956. Phloem feeder, monophagous. Specialist on native and cultivated Prunus, occasional pest on cultivated peaches and plums. Found in montane regions from the southern Rocky Mountains of the southwestern United States to Central America in pine-oak communities.
  • 98. Scolytus dimidiatus Chapuis, 1869. Phloem feeder, oligophagous? Most commonly collected in Leguminosae hosts, especially Lonchocarpus. Humid forests at intermediate and lower elevations from eastern Mexico to Venezuela.
  • 99. Hypothenemus californicus Hopkins, 1915. Pith borer, polyphagous. Known from a variety of hosts, generally herbaceous. Widely distributed but seldom collected in tropical and subtropical regions of the Americas. Southeastern US and southern California to Brazil.
  • 100. Hypothenemus crudiae (Panzer, 1791). Pith borer, polyphagous. Known from a wide variety of hosts. Eastern United States. Lowland areas in dry and humid habitats on both coasts of Mexico to Brazil and Argentina. Also found in Africa and Asia, possibly introduced there.
  • 101. Hypothenemus erectus LeConte, 1885. Pith borer,polyphagous.Known from a wide variety of hosts. Florida and southern Texas, dry and humid lowland habitats on both coasts of Mexico to Venezuela.
  • 102. Hypothenemus eruditus Westwood, 1836. Pith borer, polyphagous. Known from a wide variety of hosts. All subtropical and tropical habits in all regions of the Americas. Cosmopolitan, found on all continents with suitable habitats.
  • 103. Hypothenemus interstitialis (Hopkins, 1915). Pith borer, polyphagous. Known from a wide variety of hosts. Eastern United States. Lowland areas in dry and humid habitats on both coasts of Mexico to Brazil.
  • 104. *Hypothenemus obscurus (Fabricius, 1801). Pith borer, polyphagous. Known from a wide variety of hosts. Widely distributed in tropical and subtropical regions of the Americas.
  • 105. Hypothenemus seriatus (Eichhoff, 1872). Pith borer, polyphagous. Known from a wide variety of hosts. Widely distributed in tropical and subtropical regions of the Americas. Cosmopolitan, found on all continents with suitable habitat.
  • 106. Hypothenemus squamosus (Hopkins, 1915). Pith borer, polyphagous. Known from a wide variety of hosts. Florida and southern Texas, dry and humid lowland habitats on both coasts of Mexico and Caribbean.
  • 107. Ambrosiodmus coffeiceus (Schedl, 1951). Ambrosial, polyphagous. Known from a variety of hosts. Humid environments at intermediate elevations from eastern Mexico (Nuevo Leon) to Suriname.
  • 108. Ambrosiodmus ferus Wood, 1986. Ambrosial, monophagous? Collected only from Quercus, but other species in this genus are polyphagous. Known only from the immediate region of Xalapa.
  • 109. Ambrosiodmus rugicollis (Blandford, 1898). Ambrosial, monophagous? Collected only from Quercus, but other species in this genus are polyphagous. Intermediate elevations from eastern Mexico (Nuevo Leon) to Guatemala.
  • 110. Ambrosiodmus rusticus (Wood, 1974). Ambrosial, polyphagous. Known from a variety of hosts. Widely distributed at intermediate elevations in most mountain ranges in Mexico to Honduras.
  • 111. Coptoborus pseudotenuis (Schedl, 1936). Ambrosial, polyphagous. Known from a variety of hosts. Intermediate and lower elevations in humid forests from eastern Mexico to Peru and Brazil.
  • 112. Coptoborus vespatorius (Schedl, 1931). Ambrosial, polyphagous. Known from a variety of hosts. Intermediate and lower elevations in humid forests from eastern Mexico to Paraguay and Argentina.
  • 113. Dryocoetoides capucinus (Eichhoff, 1869). Ambrosial, polyphagous. Known from a variety of hosts. Intermediate and lower elevations in humid forests from eastern and western Mexico to Brazil.
  • 114. Euwallacea discretus (Eggers, 1933). Ambrosial, polyphagous. Known from a variety of hosts. Reported as Xyleborus discretus Eggers. Bright (2019), following Storer et al. (2015), transferred additional Neotropical species to Euwallacea. Based on the same criteria, this species also belongs here, though the combination has not been published before. Intermediate and lower elevations in humid forests from southeastern Mexico to Peru.
  • 115. Euwallacea posticus (Eichhoff, 1869). Ambrosial, polyphagous. Known from a variety of hosts. Storer et al. (2015) transferred this species to Euwallacea based on morphological and molecular data and speculated that other Neotropical species would eventually be transferred as well. Intermediate and lower elevations in humid forests from eastern Mexico to Paraguay and Argentina.
  • 116. *Theoborus theobromae Hopkins, 1915. Ambrosial, polyphagous. Known from a variety of hosts. Intermediate and lower elevations in humid forests from eastern Mexico to Brazil and Argentina.
  • 117. Xyleborinus gracilis (Eichhoff, 1868). Ambrosial, polyphagous. Known from a variety of hosts. Southeastern United States to Argentina.
  • 118. *Xyleborinus intersetosus (Blandford, 1898). Ambrosial, polyphagous. Known from a variety of hosts. Intermediate and lower elevations in humid forests from eastern Mexico to Brazil.
  • 119. Xyleborus affinis Eichhoff, 1868. Ambrosial, polyphagous. Known from a variety of hosts. Widely distributed in eastern North America and humid and dry tropical areas throughout the tropical and subtropical regions of the New World. Also found widely in Africa and Asia.
  • 120. Xyleborus bispinatus Eichhoff, 1868. Ambrosial, polyphagous. Known from a variety of hosts. This species was previously confounded with X. ferrugineus and has only recently been consistently separated (Atkinson et al. 2013). Southeastern United States to Argentina.
  • 121. Xyleborus ferrugineus (Fabricius, 1801). Ambrosial, polyphagous. Known from a variety of hosts. Found in all tropical and subtropical regions of the New World. In Mexico its distribution is limited by cold (absent from higher elevations) and low humidity (absent from desert regions). Also widely distributed in eastern North America.
  • 122. Xyleborus horridus Eichhoff, 1869. Ambrosial, polyphagous. Known from a variety of hosts. Southern Texas to Panama, lower and intermediate elevations on both coasts of Mexico.
  • 123. Xyleborus imbellis Blandford, 1898. Ambrosial, polyphagous. Known from a variety of hosts. Humid forests at intermediate and lower elevations from eastern Mexico (Tamaulipas) to Honduras and El Salvador.
  • 124. Xyleborus squamulatus Eichhoff, 1869. Ambrosial, polyphagous. Known from a variety of hosts. Humid forests at intermediate and lower elevations from eastern Mexico (Veracruz) to southern Brazil.
  • 125. Xyleborus volvulus (Fabricius, 1775). Ambrosial, polyphagous. Known from a variety of hosts. Found in all tropical and subtropical regions of the New World. In Mexico its distribution is limited by cold (absent from higher elevations) and low humidity (absent from desert regions).