Atheta auct .) picipennis (Mannerheim 1843
Creators
Description
Atheta (Dimetrota auct.) picipennis (Mannerheim, 1843)
(Figs. 3, 32 in Lohse & Smetana 1985; Figs. 63–66 in Klimaszewski & Winchester 2002)
Homalota picipennis Mannerheim, 1843: 224.
Atheta (s. str.) carlottae Casey, 1910a: 32.
Atheta (s. str.) aperta Casey, 1910a: 33, syn. nov.
Atheta (s. str.) wrangelica Casey, 1911a: 91, syn. nov.
Atheta (s. str.) morbosa Casey, 1911a: 107, syn. nov.
Atheta (s. str.) intacta Casey, 1911a: 110, syn. nov.
Atheta (s. str.) alaskana Casey, 1911a: 113, syn. nov.
Atheta (Dimetrota) picipennis: Fenyes, 1920: 204 (ex parte; as synonym of At. rugulosa (Heer, 1839)).
Atheta (s. str.) carlottae: Fenyes, 1920: 214 (as valid species).
Atheta (s. str.) aperta: Fenyes, 1920: 214 (as valid species).
Atheta (s. str.) wrangelica: Fenyes, 1920: 214 (as valid species).
Atheta (s. str.) morbosa: Fenyes, 1920: 215 (as valid species).
Atheta (s. str.) intacta: Fenyes, 1920: 215 (as valid species).
Atheta (s. str.) alaskana: Fenyes, 1920: 215 (as valid species).
Atheta (Dimetrota) picipennis: Bernhauer & Scheerpeltz, 1926: 664 (ex parte; as valid species).
Atheta (s. str.) carlottae: Bernhauer & Scheerpeltz, 1926: 639 (as valid species).
Atheta (s. str.) aperta: Bernhauer & Scheerpeltz, 1926: 638 (as valid species).
Atheta (s. str.) wrangelica: Bernhauer & Scheerpeltz, 1926: 652 (as valid species).
Atheta (s. str.) morbosa: Bernhauer & Scheerpeltz, 1926: 645 (as valid species).
Atheta (s. str.) intacta: Bernhauer & Scheerpeltz, 1926: 644 (as valid species).
Atheta (s. str.) alaskana: Bernhauer & Scheerpeltz, 1926: 638 (as valid species).
Atheta (Dimetrota) picipennis: Moore & Legner, 1975: 370 (ex parte; as valid species).
Atheta (s. str.) carlottae: Moore & Legner, 1975: 356 (as valid species).
Atheta (s. str.) aperta: Moore & Legner, 1975: 354 (as valid species; misspelled as apera).
Atheta (s. str.) wrangelica: Moore & Legner, 1975: 377 (as valid species).
Atheta (s. str.) morbosa: Moore & Legner, 1975: 366 (as valid species).
Atheta (s. str.) intacta: Moore & Legner, 1975: 364 (as valid species).
Atheta (s. str.) alaskana: Moore & Legner, 1975: 354 (as valid species).
Dimetrota picipennis: Seevers, 1978: 260 (as valid species).
Xenota carlottae: Seevers, 1978: 270 (as valid species).
Xenota aperta: Seevers, 1978: 269 (as valid species).
Xenota wrangelica: Seevers, 1978: 271 (as valid species).
Xenota morbosa: Seevers, 1978: 270 (as valid species).
Xenota intacta: Seevers, 1978: 270 (as valid species).
Xenota alaskana: Seevers, 1978: 269 (as valid species).
Dimetrota picipennis: Lohse & Smetana, 1985: 283 (as valid species).
Dimetrota carlottae: Lohse & Smetana, 1985: 283 (as synonym of Dim. picipennis).
Atheta (Dimetrota) picipennis: Klimaszewski & Winchester, 2002: 26 (as valid species).
Type material. Lectotype of Homalota picipennis (designated by Lohse and Smetana (1985)): UNITED STATES: Alaska: ♂, Sitka (MZHF).
Lectotype of Atheta carlottae (here designated): ♂, “QCI [Queen Charlotte Islands]”, “ Atheta carlottae Csy. ”, “TYPE USNM 39247” (red label), “CASEY bequest 1925” (NMNH). Paralectotypes: 2♂♂, 2♀♀, “QCI [Queen Charlotte Islands]”, “ carlottae PARATYPE USNM 39247” (red label), “CASEY bequest 1925” (NMNH).
Lectotype of Atheta aperta (here designated): ♂, “Cal.[ifornia, Humboldt Co., Arcata]”, “ Atheta aperta Csy. ”, “TYPE USNM 39250” (red label), “CASEY bequest 1925” (NMNH). Paralectotype: ♀, “Cal.[ifornia, Humboldt Co., Arcata]”, “ aperta 2 PARATYPE USNM 39250” (red label), “CASEY bequest 1925” (NMNH).
Holotype of Atheta wrangelica: ♀, “ Ft. Wrangel [Wrangell], Alaska. Wickham”, “ ♀ ”, “ wrangelica Csy. ”, “TYPE USNM 39248” (red label), “CASEY bequest 1925” (NMNH).
Lectotype of Atheta morbosa (here designated): ♀, “Cal.[ifornia, Saucelito near San Francisco]”, “ Atheta morbosa Csy. ”, “TYPE USNM 39293” (red label), “CASEY bequest 1925” (NMNH).
Holotype of Atheta intacta: ♀, “Cal.[ifornia, Marin Co., Saucelito]”, “ intacta Csy. ”, “TYPE USNM 39299” (red label), “CASEY bequest 1925” (NMNH).
Lectotype of Atheta alaskana (here designated): ♀, “Alask.[Alaska, “without further indication, but probably collected on the mainland near Fort Wrangel” (Casey 1910a)] Wick.[ham]”, “ alaskana Csy. ”, “TYPE USNM 39281” (red label), “CASEY bequest 1925” (NMNH).
Additional material: CANADA: British Columbia: ♂, Upper Carmanah Valley, 31.vii–11.viii.1991 (N.Winchester); ♀, ditto but 28.viii–9.ix.1991 (SPSU); UNITED STATES: California: San Francisco Co.: ♀ (NMNH (Casey collection)).
Diagnosis. See Lohse & Smetana (1985); Klimaszewski & Winchester (2002).
Discussion. The name Atheta picipennis (Mannerheim, 1843) (ex Homalota) is a secondary junior homonym of the name Atheta picipennis (Stephens, 1832) (ex Aleochara) which is itself a primary junior homonym of the name Aleochara picipennis Gyllenhal, 1827. Atheta picipennis (Stephens, 1832) has been considered a synonym of Atheta amicula (Stephens, 1832) since at least 1868 (Gemminger & Harold 1868) and to the best of my knowledge has not been cited as valid after 1899 (Article 23.9.1.1). In the last 50 years at least 31 works by at least 42 authors have been published (Hatch 1957; Gensicke 1960; Hansen et. al. 1960; Strand & Vik 1964; Koch 1968; Scheerpeltz 1968; Easton 1970; Palm 1970; Blackwelder 1973; Bordoni 1973; Tikhomirova 1973; Benick & Lohse 1974; Moore & Legner 1975; Yosii & Sawada 1976; Hanski & Koskela 1977; Pope 1977; Hanski & Koskela 1978; Muona 1979b; Burakowski et al. 1981; Uhlig & Vogel 1981; Benick 1982; Sawada 1982; Muona 1984; Lohse 1985; Segers 1986; Kahlen 1987; Muona 1987; Silfverberg 1988; Alexandrovitch et al. 1996; Derunkov & Melke 2001; Klimaszewski & Winchester 2002) listing Atheta picipennis as valid (Article 23.9.1.2). Atheta picipennis (Mannerheim, 1843) is a nomen protectum and Atheta picipennis (Stephens, 1832) is a nomen oblitum (Article 23.9.2).
Atheta picipennis was believed to have a Holarctic distribution (e.g., Bernhauer & Scheerpeltz 1926) until Lohse and Smetana (1985) demonstrated that the Palaearctic specimens belonged to a separate species, At. aeneipennis (Thomson, 1856).
Females of At. picipennis and At. altaica Bernhauer, 1901 have very similar spermathecae. Klimaszewski and Winchester (2002) were unable to find any difference between females of the two species. Although in average At. picipennis has a larger body than At. altaica, females of these two species can be reliably identified only by association with males. In comparison to At. altaica, males of At. picipennis have a narrower apex of the median lobe of the aedeagus.
Distribution. Atheta picipennis and At. altaica appear to be allopatric: Atheta picipennis is known from southeast Alaska (Sitka, Wrangell), coastal British Columbia and California, while At. altaica is known (in North America) from the Kenai Peninsula (Alaska), central Alaska, Yukon Territory, Northwest Territories and Labrador (Lohse et al. 1990).
Other
Published as part of Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, pp. 1-134 in Zootaxa 353 (1) on pages 40-42, DOI: 10.11646/zootaxa.353.1.1, http://zenodo.org/record/5019783Files
Files
(9.2 kB)
| Name | Size | Download all |
|---|---|---|
|
md5:a21cbdf74cb93c0b22d4a35b29914ebd
|
9.2 kB | Download |
System files
(94.0 kB)
| Name | Size | Download all |
|---|---|---|
|
md5:3561b3d8a2911bab3a55f580fce925ed
|
94.0 kB | Download |
Linked records
Additional details
Identifiers
Biodiversity
- Collection code
- MZHF , NMNH , SPSU
- Event date
- 1991-07-31 , 1991-08-28
- Family
- Staphylinidae
- Genus
- Atheta
- Kingdom
- Animalia
- Order
- Coleoptera
- Phylum
- Arthropoda
- Scientific name authorship
- Mannerheim
- Taxon rank
- genus
- Type status
- holotype , lectotype , paralectotype
- Verbatim event date
- 1991-07-31/08-11 , 1991-08-28/09-09
- Taxonomic concept label
- Atheta (Mannerheim, 1843 sec. Gusarov, 2003
References
- Mannerheim, C. G. (1843) Beitrag zur Kaefer-Fauna der Aleutischen Inseln, der Insel Sitkha. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16 (1), 175 - 314.
- Lohse, G. A. & Smetana, A. (1985) Revision of the types of species of Oxypodini and Athetini (sensu Seevers) described by Mannerheim and Maklin from North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 39 (3), 281 - 300.
- Klimaszewski, J. & Winchester, N. N. (with colour images by K. Bolte) (2002) Aleocharine rove beetles (Coleoptera Staphylinidae) of the ancient Sitka spruce forest on Vancouver Island, British Columbia, Canada. Memoires de la Societe royale belge d'Entomologie, 40, 3 - 126.
- Casey, T. L. (1910 a) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183.
- Casey, T. L. (1911 a) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, pp. 1 - 245.
- Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414.
- Heer, O. (1839) Fauna coleopterorum Helvetica. Pars I, fasc. 2. Orelii, Fuesslini et Sociorum, Turici [Zurich], pp. 145 - 360.
- Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988.
- Moore, I. & Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp.
- Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp.
- Stephens, J. F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects: con- taining their generic and specific distinctions; with an account of their metamorphoses, time of appearance, localities, food, and economy, as far as practicable. Mandibulata, 5. Baldwin and Cradock, London, 1 - 447.
- Gyllenhal, L. (1827) Insecta Suecica descripta. Classis I. Coleoptera sive Eleuterata. Tom I. Pars IV. Friederichum Fleischer, Lipsiae (Leipzig), viii + 762 pp.
- Gemminger, M. & Harold, E. von (1868) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom II. E. H. Gummi, Monachii [Munich], pp. 425 - 752 + 6 pp.
- Hatch, M. H. (1957) The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Press, Seattle, ix + 384 pp.
- Gensicke, F. (1960) Zur Kenntnis der Nestfauna einiger Muridenarten in der Umgebung von Greifswald. Wissenschaftliche Zeitschrift der Ernst-Moeritz-Arndt-Universitat Greifswald, 9, 189 - 197.
- Strand, A. & Vik, A. (1964) Die Genitalorgane der nordischen Arten der Gattung Atheta Thoms. (Col., Staphylinidae). Norsk Entomologisk Tidsskrift, 12 (5 - 8), 327 - 335, Taf. I - XXI.
- Koch, K. (1968) Kaferfauna der Rheinprovinz. Decheniana (Bonn), Beihefte 13, 1 - 382.
- Scheerpeltz, O. (1968) Coleoptera-Staphylinidae. Catalogus Faunae Austriae, 15 fa. Springer- Verlag, Wien, 279 pp.
- Easton, A. M. (1970) Atheta boletophila Thoms. (Col. Staphylinidae) new to Britain. The Entomologist's Monthly Magazine, 105 (1262 - 1264) [1969], 197 - 198.
- Palm, T. (1970) Svensk Insektfauna. 9. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Underfam. Aleocharinae (Atheta). Hafte 6. Entomologiska Foreningen i Stockholm, Stockholm, pp. 113 - 296, pl. I - XXI.
- Blackwelder, R. E. (1973) Checklist of the Staphylinidae of Canada, United States, Mexico, Central America and the West Indies. North American Beetle Fauna Project. Family no. 15, complete (Yellow Version). Biological Research Institute of America, Inc., Sienna College, Loudonville, New York, 165 pp.
- Bordoni, A. (1973) I Coleotteri Stafilinidi delle isole circumsiciliane. Lavori della Societa Italiana di Biogeografia, 3 [1972], 651 - 754.
- Tikhomirova, A. L. (1973) Morfoekologicheskiye osobennosti i filogenez stafilinid (s katalogom fauny SSSR). Nauka, Moscow, 191 pp.
- Benick, G. & Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. & Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke & Evers Verlag, Krefeld, pp. 72 - 220.
- Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140.
- Hanski, I. & Koskela, H. (1977) Niche relations among dung-inhabiting beetles. Oecologia, 28, 203 - 231.
- Pope, R. D. (1977). In: Kloet, G. S. & Hincks, W. D. (Eds) A Check List of British Insects. Second edition (completely revised). Part 3. Coleoptera and Strepsiptera. Handbooks for the Identification of British Insects, 11 (3), pp. xiv + 105.
- Hanski, I. & Koskela, H. (1978) Stability, abundance, and niche width in the beetle community inhabiting cow dung. Oikos, 31, 290 - 298.
- Muona, J. (1979 b) Staphylinidae. In: Silfverberg, H., Enumeratio Coleopterorum Fennoscandiae et Daniae. Helsingfors Entomologiska Bytesforening, Helsinki, pp. 14 - 28.
- Burakowski, B., Mroczkowski, M. & Stefanska, J. (1981) Katalog fauny Polski - Catalogus fau - nae Poloniae. Cz Osc 23, tom 8. Chrz szcze - Coleoptera. Kusakowate - Staphylinidae, cz Osc, 3 Aleocharinae. Panstwowe Widawnictwo Naukowe, Warszawa, 330 pp.
- Uhlig, M. & Vogel, J. (1981) Zur Staphylinidenfauna der Umgebung von Waren / Muritz (Mecklenburg). Unter besonderer Berucksichtigung der Naturschutzgebiete " Ostufer der Muritz ", " Ostufer des Feisnecksees ", und des Flachennaturdenkmals " Wienpietschseen ". 5. Beitrag zur Faunistik der Staphylinidae (Coleoptera). Mitteilungen aus dem Zoologischen Museum in Berlin, 57 (1), 75 - 168.
- Benick, G. (1982) Arten der Unterfamilie Athetae aus der Mongolei (Coleoptera, Staphylinidae), IV. Annales Historico-Naturales Musei Nationalis Hungarici, 74, 93 - 114.
- Sawada, K. (1982) Atheta and its allies of Southeast Asia (Coleoptera; Staphylinidae). III. Oriental species described by V. von Motschulsky and G. Kraatz. Contributions from the Biological Laboratory, Kyoto University, 26 (2), 141 - 187.
- Muona, J. (1984) Review of Palearctic Aleocharinae also occuring in North America (Coleoptera: Staphylinidae). Entomologica Scandinavica, 15, 227 - 231.
- Segers, R. (1986) Catalogus Staphylinidarum Belgicae (Coleoptera). Institut Royal des Sciences Naturelles de Belgique. Documents de Travail., No. 32, Bruxelles, 104 pp.
- Kahlen, M. (1987) Nachtrag zur Kaferfauna Tirols. Erganzung zu den bisher erschienenen faunistischen Arbeiten uber die Kafer Nordtirols (1950, 1971 und 1976) und Sudtirols (1977). Im Selbstverlag des Tiroler Landesmuseum Ferdinandeum, Innsbruck, 288 pp.
- Muona, J. (1987) Some aspects of Aleocharinae systematics - a response to Dr. G. A Lohse. Entomologische Blatter, 83 (1), 19 - 24.
- Silfverberg, H. (1988) Lists of the insect types in the Zoological Museum, University of Helsinki. 8. Coleoptera: Staphyliniade. Acta Entomologica Fennica, 52, 15 - 40.
- Derunkov, A. V. & Melke, A. (2001) Familia Staphylinidae. In: Gutowski, J. M. & Jaroszewicz, B. (Eds.), Catalogue of the fauna of Bia 1 owie Z a Primeval Forest. IBL, Warszawa, pp. 133 - 147.
- Thomson, C. G. (1856) Nagra nya arter af Insekt-slagtet Homalota. Ofversigt af Kongl. Vetenskaps- Akademiens Forhandlingar, 13 (4), 91 - 107.
- Bernhauer, M. (1901) Neue Staphyliniden aus Centralasien. Verhandlungen der k. k. zoologischbotanischen Gesellschaft in Wien, 51, 106 - 115.
- Lohse, G. A., Klimaszewski, J. & Smetana, A. (1990) Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 44 (2), 121 - 202.