Anatoma jansenae Geiger, 2006, new species

Anatoma jansenae new species: Figure 18

Anatoma SWA: Jansen 1999: 50, figs. 16–18.

Anatoma australis partim: Geiger & Jansen 2004a: fig. 5 (A. jansenae), not figs. 3–4 (A. australis).

Type material. Holotype (AMS C.402717). Paratypes (AMS C.450272, 7; AMS C.402720, 7; AMS C.402721, 1).

Type locality. 238– 183 m, SW of Cape Naturaliste, Western Australia, Australia, 33.742°S, 114.435°E.

Etymology. Named after Patty Jansen, New South Wales, Australia, who first recognized the distinctness of the species, and for her contributions to the knowledge of Australian micromolluscs.

Description. Shell trochiform globular, medium size (to 2.5 mm). Protoconch of 0.75 whorls, with flocculant sculpture, apertural varix not connected to embryonic cap, apertural margin somewhat sinusoid. Teleoconch I of 0.75 whols, 17–21 distinct axials, spiral of same strength as axial in position of selenizone. Teleoconch II of up to 2.25 whorls. Shoulder somewhat convex, approximately 75 distinct axials on last whorl, 12–18 fine spirals forming at intersection with axials minute, dull points. Base with similar sculpture as shoulder, approximately 20 somewhat stronger spirals; spirals always cords, not as shingles. Umbilicus narrow, continuously sloping with base. Selenizone at periphery, keels moderately strong, moderately elevated; slit open margins converging towards apertural rim. Aperture rounded in lower portion, suborbicular under shoulder.

Differential diagnosis. Anatoma australis (Hedley, 1903: Fig. 19) from the eastern Australia has coarser sculpture on the base, particularly with a decrease in density of the spirals in the third closest to the umbilicus; adumbilical margin of adumbilical spirals usually not fully differentiated from surface giving shingle­like appearance. Anatoma tobeyoides Geiger & Jansen, 2004 from southeastern Australia lacks the protoconch varix, and on teleoconch I of less than 0.5 whorls lacks a spiral cord in the position of the selenizone. Anatoma funiculata Geiger & Jansen, 2004 from Queensland is less globular and more angular in overall shape, has a teleoconch I of approximately 0.5 whorls, and a distinct shell strand (funiculus) running into the umbilicus.

Distribution. Western Australia, Coral Sea.

Specimen records. Coral Sea. 6 m, Saumarez Reef, 21.817°S, 153.667°E (AMS C.402669, 1).

Timor Sea. 27 m, Sahul Banks, 11.5°S, 125.5°E (AMS C.377600, 1).

Australia, Western Australia. 75 m, Off Albany, 35.240°S, 118.342°E (AMS C.378550, 7). 158 m, Great Australian Bight, E of Hood Point, 34.417°S, 121.333°E (AMS C.378551, 2). South Cowaramup, 33.883°S, 114.983°E (AMS C.379071, 7). 238– 183 m, SW of Cape Naturaliste, 33.742°S, 114.435°E (AMS C.402717, 1: holotype; AMS C.ex. C.402717, 7: paratypes). 155 m, NW of Bunbury, 33.250°S, 114.617°E (AMS C.402719, 1). 200–221 m, NW of Bunbury, 33.000°S, 114.617°E (AMS C.402747, 6). 176–182 m, W of Garden Island, 32.262°S, 115.112°E (AMS C.402741, 1). 210–212 m, W of Garden Island, 32.250°S, 115.117°E (AMS C.402750, 2). Minim Cove, Mosman Park, Swan River, 32.017°S, 115.767°E (AMS C.379070, 1). 116 m, off Rottnest Island, 31.673°S, 115.198°E (AMS C.378552, 8). 160 m, off Rottnest Island, 31.650°S, 115.080°E (AMS C.402749, 1). 110 m, off Rottnest Island, 31.630°S, 115.178°E (AMS C.378549, 3). 732 m, W of Rottnest Island, 31.083°S, 114.767°E (AMS C.402751, 1). 237–274 m, W of Green Head, 30.750°S, 114.767°E (AMS C.402748, 1). 238–247 m, off Cervantes Island, 30.533°S, 114.683°E (AMS C.402752, 2). 256– 192 m, NW of Cervantes, 30.500°S, 114.633°E (AMS C.402718, 10). 223–245 m, off Jurien Bay, 30.133°S, 114.500°E (AMS C.402745, 1). 197–219 m, NW of Green Head, 29.967°S, 114.450°E (AMS C.402721, 1: paratype). 183 m, NW of Beagle Island, 29.725°S, 114.333°E (AMS C.402746, 1). 274–283 m, NW of Beagle Island, 29.717°S, 114.283°E (AMS C.402720, 7: paratypes). 146 m, W of Dongara, 29.350°S, 114.117°E (AMS C.402742, 1). 219 m, W.of Dongara, 29.183°S, 113.900°E (AMS C.402744, 1). 183 m, W of Dongara, 29.142°S, 113.913°E (AMS C.402743, 1). Point Quobba, N of Carnarvon, 24.483°S, 113.417°E (AMS C.379976, 1). 108 m, Off North West Cape, 22.705°S, 113.540°E (AMS C.402644, 1; 402645, 1). 2 m, Ningaloo Reef, off Neds Camp, 21.992°S, 113.908°E (AMS C.377284, 2). 238 m, North West Shelf, ca 230 ml W Roebuck Bay, 18.500°S, 118.050°E (AMS C.402676, 1).

Remarks. Geiger & Jansen (2004) noted that there was no consistent difference in the eastern and western specimens of A. australis and Anatoma “SWA” of Jansen (1999).

Recent re–examination of the material showed the above indicated subtle differences.

Anatoma munieri (Fischer, 1862): Figure 20

Scissurella munieri Fischer October 1, 1862: 390–391, not illustrated.

Scissurella munieri: Munier Chalmas 1865: 397.

Scissurella munieri: Fischer 1867: 305, 468, pl,. 9, fig. 4 [The figure caption on plate 9 for figure 4 reads Scissurella munieriana, an error noted on the errata page 468 of the volume].

Scissurella munieri: Paetel 1888: 289.

Scissurella munieri: Pilsbry 1890: 54.

Scissurella munieri: Thiele 1912: 14 –15.

scissurellid: Bandel 1991: pl. 2, fig. 7.

Scissurella munieri: Higo & Goto 1993: 15.

Scissurella ? munieri: Geiger 2003: 77.

Synonyms

+ Anatomus turbinatus A. Adams November, 1862: 347 –348, not illustrated. Type material. Holotype (BMNH 1874.5.19.62. Higo et al. 2001: G82), 2.5 mm. Type locality. Mino­sima, [= Mishima Island, Hagi City, Yamaguchi Prefecture, Japan (34.767°N, 131.166E): see also Kawamoto and Tanabe (1956)] 63 fms. [= 115 m] (OD). Etymology. Turbinatus: Latin adjective: that which whirls; whirlwind, tornado; spinning top; spiral. Referring to the high spired shell shape.

Scissurella turbinata: Crosse 1863: 109.

Scissurella turbinata: Paetel 1888: 289.

Anatomus turbinatus: Pilsbry 1890: 59.

Scissurella turbinata: Pilsbry 1895: 106.

Scissurella turbinata: Thiele 1912: 15, pl. 2, figs. 9–10 [after a specimen in the British Museum: type?].

Schizotrochus turbinatus: Habe 1951: 68, pl. 11, figs. 12–13.

Scissurella turbinata: Kuroda & Habe 1952: 85.

Scissurella (Schizotrochus) turbinata: Kawamoto & Tanabe 1956: 3, pl. 2, fig. 11 [copy figure Thiele 1912. fide T. Sasaki pers. comm. 2005].

Anatoma turbinata: Habe & Kosuge, 1964: 4.

Anatoma turbinata: Higo 1973: 13.

Anatoma turbinata: Tsuchida et al. 1991: 5 –6, pl. 1, figs. 2–3, 5.

Anatoma turbinata: Higo & Goto 1993: 15.

Anatoma turbinata: Yu &Feng 1996: pl. 1,figs. 1–4.

Anatoma turbinata: Okutani & Hasegawa 2000: 37, fig. 5.

Anatoma turbinata: Higo et al. 2001: G82 [holotype].

Anatoma turbinata: Geiger 2003: 74.

Anatoma turbinata: Geiger & Jansen 2004a: 18 –21. figs. 9–10, 18 [map].

Anatoma turbinata: Geiger 2004: text­fig. p. 5.

Misidentifications

Anatoma agulhasensis: Bandel 1998: 34 –35, pl. 11, figs. 4–6. [is A. munieri].

Anatoma agulhasensis: Jansen 1999: 48,figs. 1–3. [is A. munieri].

not Scissurella turbinata: Yokoyama (1924: 35–36, pl. 5, fig. 21) [is Sci. staminea. Fide Oyama 1973: 10. plate reprinted in Taki & Oyama 1954: pl. 42: shows Sci. staminea. Specimen in UMUT CM21891 fide T. Sasaki pers. comm. 2005).

Type material. Syntypes (MNHN, 2), 1.5 x 1.66 mm. Lectotype here designated (see remarks).

Type locality. Seas of China, in bottom sands (OD).

Etymology. Named after E. Munier Chalmas (OD).

Description and differential diagnosis. The species was recently treated by Geiger & Jansen (2004a) as A. turbinata.

Remarks. Anatoma munieri and A. turbinata are clearly synonymous. Supporting characters include the overall turreted shape of the shell, the strong constriction below the selenizone with a subsequent spiral edge, a minor spiral edge on the base 0.66 towards the umbilicus, the shell ornamentation showing a regular reticulate pattern composed of spiral and axial cord, and the open umbilicus, which slopes continuously with the base.

Anatoma munieri has only one month priority over A. turbinata. The date on the first page of the issue in which Sci. munieri was described, agrees with the publication dates given by Winckworth (1936). Although the latter species name is somewhat better known, the general rule of priority should be enforced in this case. It is not possible to apply the nomen oblitum/protectum rules, because munieri has been used as a valid species after 1900.

Fischer­Piette (1950: 69) indicated there to be a holotype and one paratype. The holotype was not specifically designated in the original description, hence, the two specimens constitute syntypes. The subsequent holotype indication can not be viewed as a lectotype designation (ICZN Art. 74.5). The specimen shown in Figure 20 A is here designated as the lectotype with the express purpose of taxon stabilization in case other non­conspecific syntypes should be located.

Bandel (1991) showed an unidentified “scissurellid”, which is here identified as A. munieri. Bandel (1998: 42) referred in his discussion of Hainella pulchella (= A. pulchella: see Geiger, 2003 for discussion of generic taxa) to an illustration in his 1991 publication that illustrated the species with a specimen from Cebu. There was no specific reference to pagination, plate or figure number, and three Bandel (1991) references were used, but only one dealing with specimens from the Philippines: Bandel (1991) of this contribution. Although A. pulchella is very distinct from A. munieri, it is likely that the reference by Bandel (1998) was a lapsus calami.