Rhamphomyia (Pararhamphomyia) kjellmanii Holmgren

Rhamphomyia (Pararhamphomyia) kjellmanii Holmgren

(Fig. 35)

Rhamphomyia kjellmanii Holmgren, 1880: 22. Type locality (by lectotype designation): Novaya Zemlya, Arkhangelskaya Province, Russia.

Other references: Holmgren & Aurivillius, 1883: 163 (expedition results); Bezzi, 1903: 226 (catalogue); Melander, 1928: 195 (catalogue).

Rhamphomyia kjellmani Holmgren: Frey, 1915: 3, 8 (faunistic survey).

Rhamphomyia (Pararhamphomyia) kjellmanii Holmgren: Frey, 1955b: 500 (revision); Chvála & Wagner, 1989: 301 (catalogue); Yang et al., 2007: 172 (catalogue); Shamshev, 2016: 64 (checklist); Shamshev & Sinclair, 2018: 311 (lectotype designation, redescription); Sinclair et al., 2019: 54 (revision).

Rhamphomyia uralensis Becker, 1915: 58. Type-locality (by lectotype designation): mouth of Kara River (about 69°02′N 64°35′E), Yamalo-Nenets, Russia.

Rhamphomyia (Pararhamphomyia) uralensis Becker: Frey, 1955b: 500 (revision); Chvála & Wagner, 1989:306 (catalogue); Yang et al., 2007: 178 (catalogue); Shamshev, 2016: 71 (checklist); Sinclair et al., 2019: 54 (synonym, lectotype designation).

Material examined. RUSSIA. Archangelskaya Prov. (Novaya Zemlya Archipelago): steamer deck, in sea near western shore of Novaya Zemlya, 20.vii.1948, Korotkevich (1 ♂, 1 ♀, ZIN). Chukotka AO (Wrangel Island) : middle flow of Mamontovaya River, 71°10′N 179°45′W, BT 20 A, Sw, 2.vii.2015, OAK (1 ♂, ZIN); same locality and biotope, YPT, 2–4.vii.2015, OAK (3 ♂, ZIN); same locality, BT 3, Sw, 1.vii.2015, OAK (3 ♂, ZIN); same locality, BT 2, YPT, 2–4.vii.2015, OAK (1 ♂, ZIN); same locality, BT 1 A, Sw, 2.vii.2015, OAK (1 ♂, ZIN); same locality and biotope, YPT, 2–4.vii.2015, OAK (1 ♂, 2 ♀, ZIN); same locality, BT 7, Sw, 1.vii.2015, OAK (1 ♂, ZIN); upper flow of Neizvestnaya River, 71°13′N, 179°19′W, BT 12, Sw, 6.vii.2015, OAK (7 ♂, ZIN); same locality, BT 13, YPT, 6–15.vii.2015, OAK (1 ♂, ZIN); same locality, BT 7 A, PT, 25.vi.–10.vii.2015, OAK (1 ♂, ZIN); same locality, BT C1, Sw, 24.vi.2015, OAK (1 ♂, ZIN). Chukotka AO (Kolyuchin Island) : 25–27.vii.1938, Hecker (4 ♂, 8 ♀, ZIN).

Recognition. Mid-sized (wing length 4.2–4.5 mm) species. Male (Fig. 35) holoptic; scutum densely brownish grey pruinescent, dc uniserial, acr biserial, notopleuron with 4–6 long strong blunt-tipped setae, 6 sctl, laterotergite with pale setae; legs dark brown, shiny, fore coxa with strong blunt-tipped setae anteriorly, hind tibia clavate and clothed in dense long hair-like setae, hind tarsomeres 1–2 thickened, with similar setae dorsally; wing faintly infuscate, CuA+CuP incomplete, halter yellowish (sometimes dirty yellow); abdomen faintly greyish pruinescent, covered with pale setae; terminalia: cercus short, one-third length of epandrium, tapered apically; subepandrial lobe present, longer than cercus, clavate; epandrium gradually tapered, bearing numerous long, pale setae apically; phallus well exposed, slender, gently sinuate on about middle, looping beyond epandrium and curving back to cercus. Female wing faintly brownish infuscate, hind tibia and tarsus slender, legs with simple short setae.

Distribution. Holarctic; in Eurasia, R. kjellmanii extends across arctic and subarctic Russia, including both continental areas (Taymyr, Yakutia, Chukotka) and islands (Novaya Zemlya, Wrangel, Kolyuchin). In North America, this species is known only from the extreme northwestern region (Sinclair et al. 2019).

Habitat. On Wrangel Island this species was collected mainly in wet and, less commonly, in moderately moistened habitats in the warmest central part of the island.