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Review of ants (Hymenoptera: Formicidae) of Crete, with keys to species determination and zoogeographical remarks

Creators

  • 1. Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA; email: sdsalata@gmail.com
  • 2. Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51-148 Wrocław, Poland; e-mail: lech.borowiec@uwr.edu.pl
  • 3. Natural History Museum of Crete, University of Crete, Greece; e-mail: atrichas@nhmc.uoc.gr

Description

We present the first critical review of all Formicidae species found on Crete. Samples were collected from 302 localities, dispersed on the whole island (Prefectures of Chania – 75, Heraklion – 86, Lasithi – 74 and Rethymnon – 67). As a result, we present a detailed catalogue of 100 Cretan ant species, representatives of 6 subfamilies and 28 genera, with keys to their determination. The most diverse genera are Temnothorax (17 species), Camponotus (11) and Tetramorium (9). Additionally, we provide the very first documentation of Tapinoma festae male genitalia. All species and keys to their determination are illustrated, short notes on their biology, as well as maps on their distribution on the island, are also given.
Among 18 known endemic species 89% belong to the Myrmicinae subfamily. With 8 endemic species the Temnothorax genus is the most diverse in this group. Among all species recorded from Crete we could distinguish 14 chorotypes. The dominant groups are chorotypes of species more or less widely spread in the Mediterranean countries. They represent 59% of known ant fauna of Crete. Among them the dominant are Mediterranean (23%), Aegaean (13%) and NE-Mediterranean (9%). Nevertheless, alarming is also a high percentage of invasive taxa recorded from the island (10%).

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References

  • Agosti D. 1990. Review and reclassification of Cataglyphis (Hymenoptera, Formicidae). Journal of Natural History 24: 1457–1505.
  • Agosti D., Collingwood C.A. 1987a. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 60: 51–62.
  • Agosti D., Collingwood C.A. 1987b. A provisional list of the Balkan ants (Hym. Formicidae) with a key to the worker caste. II. Key to the worker caste, including the European species without the Iberian. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 60: 261–293.
  • Aguayo C.G. 1932. Notes on West Indian ants. Bulletin of the Brooklyn Entomological Society 27: 215–227.
  • André Ern. 1881. Catalogue raisonné des Formicides provenant du voyage en Orient de M. Abeille de Perrin et description des espèces nouvelles. Annales de la Société Entomologique de France (6)1: 53–78.
  • André Ern. 1883. Les fourmis. [concl.]. Pp. 345–404, In: André Edm. 1883. 1881–1886, Species des Hyménoptères d'Europe et d'Algérie. Tome Deuxième. Beaune: Edmond André, 919 + 48 ss.
  • Arnol'di K.V. 1928. Studien über die Systematik der Ameisen. II. Stenamma Westw. Zoologischer Anzeiger 75: 199–215.
  • Arnol'di K.V. 1930. Studien über die Systematik der Ameisen. V. Der erste Vertreter der Tribe Proceratiini (Formicidae) in USSR. Zoologischer Anzeiger 91: 143–146.
  • Arnol'di K.V. 1933. Formicidae – ants. [In Russian.]. Pp. 594–605, In: Filip'ev I.N., Ogloblin D.A. (Ed.) 1933. Keys to insects. [In Russian]. Moskwa. Ogiz, viii + 820 pp.
  • Arnol'di K.V. 1948. Ants of Talysh and the Diabar depression. Their importance for the characterization of communities of terrestrial invertebrates and for historical analysis of the fauna. [In Russian]. Trudy Zoologicheskogo Instituta. Akademiya Nauk SSSR 7(2): 206–262.
  • Arnol'di K.V. 1964. Higher and specialized representatives of the ant genus Cataglyphis (Hymenoptera, Formicidae) in the fauna of the USSR. [In Russian]. Zoologicheskii Zhurnal 43: 1800–1815.
  • Baroni Urbani C. 1964. Su alcune formiche raccolte in Turchia. Annuario dell'Istituto e Museo di Zoologia dell'Università di Napoli 16: 1–12.
  • Baroni Urbani C. 1969. Gli Strongylognathus del gruppo huberi nell'Europa occidentale: saggio di una revisione basata sulla casta operaia (Hymenoptera Formicidae). Bollettino della Società Entomologica Italiana 99–101: 132–168.
  • Baroni Urbani C. 1971. Studien zur Ameisenfauna Italiens XI. Die Ameisen des Toskanischen Archipels. Betrachtungen zur Herkunft der Inselfaunen. Revue Suisse de Zoologie 78: 1037–1067.
  • Baroni Urbani C. 1974. Studi sulla mirmecofauna d'Italia. XII. Le Isole Pontine. Fragmenta Entomologica 9: 225–252.
  • Beerli P., Hotz H., Uzzell T. 1996. Geologically dated sea barriers calibrate a protein clock for Aegean water frogs. Evolution 50: 1676–1687.
  • Begdon J. 1932 ("1931"). Wymiary i wskazniki niektórych znamion mrówki Stenamma Westw. westwoodi Arn. (Westw?) polonicum nov. subsp., znalezionej na Pomorzu. Sprawozdania Komisji Fizjograficznej oraz Materjaly do Fizjografji Kraju 65: 113–119.
  • Bergmeier E., Kypriotakis Z., Jahn R., Böehling N., Dimopoulos P., Raus T., Tzanoudakis D. 2001. Flora and phytogeographical significance of the islands Chrisi, Koufonisi and nearby islets (S. Aegean, Greece). Wildenowia 31: 329–356.
  • Bernard F. 1950 ("1946"). Notes sur les fourmis de France. II. Peuplement des montagnes méridionales. Annales de la Société Entomologique de France 115: 1–36.
  • Bernard F. 1953 ("1952"). La réserve naturelle intégrale du Mt Nimba. XI. Hyménoptères Formicidae. Mémoires de l'Institut Français d'Afrique Noire 19: 165–270.
  • Bernard F. 1956. Révision des fourmis paléarctiques du genre Cardiocondyla Emery. Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 47: 299–306.
  • Bernard F. 1957. Xenometra Emery, genre de fourmis parasite nouveau pour l'Ancien Monde (Hym. Formicidae). Bulletin de la Société Entomologique de France 62: 100–103.
  • Bernard F. 1978. Orthocrema esterelana, espèce nouvelle commune dans l'Estérel (Hym. Formicidae). Bulletin de la Société Entomologique de France 83: 43–46.
  • Blondel J, Aronson J, Bodiou J-Y, Boeuf G. 2010. The Mediterranean Region. Biological Diversity in Space and Time. Oxford University Press Inc., New York, NY.
  • Boer P. 2013. Revision of the European ants of the Aphaenogaster testaceopilosa–group (Hymenoptera: Formicidae). Tijdschrift voor entomologie 156(1): 57–93.
  • Bolton B. 1987. A review of the Solenopsis genus–group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History), Entomology 54(3): 263–452.
  • Bolton B. 1995. A new general catalogue of the ants of the world. Cambridge, Mass. Harvard University Press, 504 ss.
  • Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (monograph) 25: 1–340.
  • Borowiec L., Galkowski C., Salata S. 2015. What is Tetramorium semilaeve André, 1883? (Hymenoptera, Formicidae). ZooKeys 512: 39–62.
  • Borowiec L., Salata S. 2012. Ants of Greece – checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23: 461–563.
  • Borowiec L., Salata S. 2013. Ants of Greece – additions and corrections (Hymenoptera: Formicidae). Genus 24: 335–401.
  • Borowiec L., Salata S. 2014a. Redescription of Camponotus nitidescens Forel, 1889, new status and notes on ants from Kefalonia, Greece (Hymenoptera: Formicidae). Genus 25: 499–517.
  • Borowiec L., Salata S. 2014b. Review of Mediterranean members of the Aphaenogaster cecconii group (Hymenoptera: Formicidae), with description of four new species. Zootaxa 3861(1): 40–60.
  • Borowiec L., Salata S. 2017a. New records of ants (Hymenoptera: Formicidae) from Sterea Ellas, Greece. Acta entomologica silesiana 25(020): 143–145.
  • Borowiec L., Salata S. 2017b. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193–235.
  • Borowiec L., Salata S. 2018a. New records of ants (Hymenoptera: Formicidae) from Epirus, Greece. Acta entomologica silesiana 26(001): 1–22.
  • Borowiec L., Salata S. 2018b. Ants from Thessaly, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 87: 217–248.
  • Borowiec L., Salata S. 2018c. Notes on ants (Hymenoptera: Formicidae) of Zakynthos Island, Greece. Annals of the Upper Silesian Museum in Bytom, Entomology 27(004): 1–13.
  • Borowiec L., Salata S. 2018d. Notes on ants (Hymenoptera: Formicidae) of the Euboea Island, Central Greece. Annals of the Upper Silesian Museum in Bytom, Entomology 27(005): 1–15.
  • Bračko G. 2006. Review of the ant fauna (Hymenoptera: Formicidae) of Croatia. Acta Entomologica Slovenica 14: 131–156.
  • Bračko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15–24.
  • Bračko G., Wagner H.C., Schulz A., Gioahin E., Matičič J., Tratnik A. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North–Western Journal of Zoology 10(1): 10–24.
  • Bračko G., Kiran K., Karaman C., Salata S., Borowiec L. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945.
  • Brèthes J. 1914. Note sur quelques Dolichodérines argentines. Anales del Museo Nacional de Historia Natural de Buenos Aires 26: 93–96.
  • Brown W.L., Wilson E.O. 1957. A new parasitic ant of the genus Monomorium from Alabama, with a consideration of the status of genus Epixenus Emery. Entomological News 68: 239–246.
  • Brullé G.A. 1833 ("1832"). Expédition scientifique de Morée. Section des sciences physiques. Tome III. Partie 1. Zoologie. Deuxième section - Des animaux articulés. [part]. Paris: Levrault, pp. 289–336.
  • Buschinger A. 1989. Workerless Epimyrma kraussei Emery, 1915, the first parasitic ant of Crete. Psyche 96(1–2): 69–74.
  • Buschinger A., Douwes P. 1993. Socially parasitic ants of Greece. Biologia Gallo-Hellenica 20: 183–189.
  • Chopard L. 1921. La fourmi d'Argentine Iridomyrmex humilis var. arrogans Santschi dans le midi de la France. Annales des Epiphyties 7: 237–265.
  • Collingwood C.A., Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna of Saudi Arabia 15: 300–385.
  • Costa A. 1884. Notizie ed osservazioni sulla geo-fauna Sarda. Memoria terza. Risultamento delle ricerche fatte in Sardegna nella estate del 1883. Atti della Reale Accademia delle Scienze Fisiche e Matematiche. Napoli 1(9): 1–64.
  • Csősz S., Heinze J., Mikó I. 2015. Taxonomic Synopsis of the Ponto-Mediterranean Ants of Temnothorax nylanderi Species-Group. PloS One 10(11): e0140000.
  • Csősz S., Salata S., Borowiec L. 2018. Three Turano-European species of the Temnothorax interruptus group (Hymenoptera: Formicidae) demonstrated by quantitative morphology. Myrmecological News 26:101-119.
  • Csősz S., Schulz A. 2010. A taxonomic review of the Palaearctic Tetramorium ferox species–complex (Hymenoptera, Formicidae). Zootaxa 2401: 1–29.
  • Czechowski W., Radchenko A., Czechowska W., Vepsäläinen K. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 496 pp.
  • Dalla Torre C. 1893. Catalogus Hymenopterorum, husque descriptorum systematicus et synonymicus. Lipsiae 7: 289 pp.
  • Dermitzakis D.M. 1990. Paleogeography, geodynamic processes and event stratigraphy during the Late Cenozoic of the Aegean area. Atti Convegni Lincei 85: 263–288.
  • di Castri F. 1981. Mediterranean–type shrublands of the world, In: di Castri F, Goodall DW, Specht RL (Eds.), Mediterranean-type shrublands of the world ecosystems of the World II: Mediterranean-type Shrublands. Elsevier, Amsterdam, pp. 1–52.
  • Dlussky G.M. 1967. Ants of the genus Formica (Hymenoptera, Formicidae, g. Formica). [In Russian]. Moskva: Nauka Publishing House, 236 pp.
  • Donisthorpe H. 1950. Two more new ants from Turkey. Entomologist's Record and Journal of Variation 62: 68–69.
  • Emery C. 1869. Enumerazione dei formicidi che rinvengonsi nei contorni di Napoli con descrizioni di specie nuove o meno conosciute. Annali dell'Accademia degli Aspiranti Naturalisti. Secunda Era 2: 1–26.
  • Emery C. 1875 ("1876"). Ueber hypogaeische Ameisen. Stettiner Entomologische Zeitung 37: 71–76.
  • Emery C. 1877. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte prima. Formiche provenienti dal Viaggio dei signori Antinori, Beccari e Issel nel Mar Rosso e nel paese dei Bogos. [part]. Annali del Museo Civico di Storia Naturale 9: 363–368.
  • Emery C. 1878. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte seconda. Formiche dell'Europa e delle regioni limitrofe in Africa e in Asia. [part]. Annali del Museo Civico di Storia Naturale 12: 43–48.
  • Emery C. 1881. Viaggio ad Assab nel Mar Rosso dei Signori G. Doria ed O. Beccari con il R. Avviso "Esploratore" dal 16 novembre 1879 al 26 febbraio 1880. I. Formiche. Annali del Museo Civico di Storia Naturale 16: 525–535.
  • Emery C. 1894a. Alcune formiche dell'isola di Crete. Bullettino della Società Entomologica Italiana 26: 7–10.
  • Emery C. 1894b. Studi sulle formiche della fauna neotropica. VI-XVI. Bullettino della Società Entomologica Italiana 26: 137–241.
  • Emery C. 1895a. Sopra alcune formiche della fauna mediterranea. Memorie della Reale Accademia delle Scienze dell'Istituto di Bologna 5: 59–75.
  • Emery C. 1895b. Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 8: 685–778.
  • Emery C. 1898. Beiträge zur Kenntniss der palaearktischen Ameisen. Öfversigt af Finska Vetenskaps-Societetens Förhandlingar 20: 124–151.
  • Emery C. 1906. Rassegna critica delle specie paleartiche del genere Myrmecocystus. Memorie della Reale Accademia delle Scienze dell'Istituto di Bologna (6)3: 47–61.
  • Emery C. 1908a. Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. II. Camponotus der herculaenus–und maculatus–Gruppe. Deutsche Entomologische Zeitschrift 6: 182–205.
  • Emery C. 1908b. Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. (Hym.) III. Die mit Aphaenogaster verwandte Gattungengruppe. Deutsche Entomologische Zeitschrift 6: 305–338.
  • Emery C. 1908c. Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. (Hym.) Teil III. Deutsche Entomologische Zeitschrift 6: 437–465.
  • Emery C. 1908d. Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. (Hym.) IV. Parasitische und Gast–Myrmicinen mit Ausnahme von Strongylognathus. Deutsche Entomologische Zeitschrift 6: 549–558.
  • Emery C. 1908e. Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. (Hym.) Teil V. Deutsche Entomologische Zeitschrift 6: 663–686.
  • Emery C. 1908f. In: Cecconi G. 1908. Contributo alla fauna delle Isole Tremiti. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Università di Torino 23(583): 1–53.
  • Emery C. 1912. Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. Teil XI. Deutsche Entomologische Zeitschrift 10: 651–672.
  • Emery C. 1914. Wissenschaftliche Ergebnisse der Bearbeitung von O. Leonhard's Sammlungen. 5. Südeuropäische Ameisen (Hym.). Entomologische Mitteilungen. Berlin-Dahlem 3: 156–159.
  • Emery C. 1915a. Contributo alla conoscenza delle formiche delle isole italiane. Descrizioni di forme mediterrane nuove o critiche. Annali del Museo civico di storia naturale di Genova 6(46)(1–27): 244–270.
  • Emery C. 1915b. Escursioni zoologiche del Dr. Enrico Festa nell'Isola di Rodi. XII. Formiche. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Università di Torino 30(701): 1–7.
  • Emery C. 1915c. Les Pheidole du groupe megacephala (Formicidae). Revue Zoologique Africaine (Brussels) 4: 223–250.
  • Emery C. 1916 ("1915"). Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79–275.
  • Emery C. 1921a. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. In: Wytsman P. (Ed.), Genera insectorum. Fasc. 174a, pp. 1–94
  • Emery C. 1921b. Formiche raccolte a Budrum (Anatolia) da Raffaele Varriale, Cap. medico nella R. Marina. Annali del Museo Civico di Storia Naturale "Giacomo Doria" 49[=(3)9]: 208–218.
  • Emery C. 1921c. Notes critiques de myrmécologie. [IX-X.]. Annales de la Société Entomologique de Belgique 61: 313–319.
  • Emery C. 1922. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. In: Wytsman P. (Ed.), Genera insectorum. Fasc. 174b, pp. 95–206.
  • Emery C. 1924a. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [concl.], Genera insectorum. Fasc. 174c, pp. 207–397.
  • Emery C. 1924b. Formiche di Spagna raccolte dal Prof. Filippo Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 17: 164–171.
  • Emery C. 1925a. Révision des espèces paléarctiques du genre Tapinoma. Revue Suisse de Zoologie 32(2): 45–64.
  • Emery C. 1925b. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183: 1–302.
  • Fabricius J.C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species adiectis synonymis, locis, descriptionibus, observationibus. Flensburgi et Lipsiae: Korte, 832 pp.
  • Fabricius J.C. 1787. Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus, genericis, differentiis, specificis, emendationibus, observationibus. Tome I. Hafniae: C. G. Proft, 348 pp.
  • Fabricius J.C. 1793. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species, adjectis synonimis, locis observationibus, descriptionibus. Tome 2. Hafniae: C. G. Proft, 519 pp.
  • Fabricius J.C. 1804. Systema Piezatorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Brunswick: C. Reichard, xiv + 15–439 + 30 pp.
  • Fattorini S. 2000. Dispersal, vicariance and refuges in the Anatolian Pimeliinae (Coleoptera, Tenebrionidae): remarks on some biogeographical tenets. Biogeographia 21: 355–398.
  • Finzi B. 1922 ("1921"). Primo contributo alla conoscenza della fauna mirmecologica della Venezia Giulia. Bullettino della Società Entomologica Italiana 53: 118–120.
  • Finzi B. 1929. Le forme italiane del genere Messor. Bollettino della Società Entomologica Italiana 61: 75–94.
  • Finzi B. 1930a. Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes. XII. Die Ameisen der Jonischen Inseln. Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I, 139(5–6): 309–319.
  • Finzi B. 1930b. Hymenopteren aus Palästina und Syrien. (Zoologische Studienreise von R. Ebner 1928 mit Unterstützung der Akademie der Wissenschaften in Wien.) Formicidae. Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I, 139: 22–24.
  • Finzi B. 1939. Ergebnisse der von Franz Werner und Otto v. Wettstein auf den Ägäischen Inseln unternommenen Sammelreisen. Ameisen. Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I, 148: 153–161.
  • Foerster A. 1850a. Hymenopterologische Studien. 1. Formicariae. Aachen: Ernst Ter Meer, 74 pp.
  • Foerster A. 1850b. Eine Centurie neuer Hymenopteren. Zweite Dekade. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalenss 7: 485–500.
  • Fonscolombe B. de 1846. Notes sur huit espèces nouvelles d'Hyménoptères et de Neuroptères, trouvées aux environs d'Aix. Annales de la Société Entomologique de France (2)4: 39–51.
  • Forel A. 1874. Les fourmis de la Suisse. Systématique, notices anatomiques et physiologiques, architecture, distribution géographique, nouvelles expériences et observations de moeurs. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft für die Gesammten Naturwissenschaften 26: 1–452.
  • Forel A. 1878. Études myrmécologiques en 1878 (première partie) avec l'anatomie du gésier des fourmis. Bulletin de la Société Vaudoise des Sciences Naturelles 15: 337–392.
  • Forel A. 1886. Nouvelles fourmis de Grèce récoltées par M. E. von Oertzen. Annales de la Société Entomologique de Belgique 30: CLIX–CLXVIII.
  • Forel A. 1889 (1888). Ameisen aus den Sporaden, den Cykladen und Griechenland gesammelt 1887 von Herrn v. Oertzen. Berliner Entomologischer Zeitschrift 32(2): 255–265.
  • Forel A. 1890. Fourmis de Tunisie et de l'Algérie orientale. Annales de la Société Entomologique de Belgique 34: lxi–lxxvi.
  • Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 42: 305–318.
  • Forel A. 1893. Nouvelles fourmis d'Australie et des Canaries. Annales de la Société Entomologique de Belgique 37: 454–466.
  • Forel A. 1894a. Les Formicides de la Province d'Oran (Algérie). Bulletin de la Société Vaudoise des Sciences Naturelles 30: 1–45.
  • Forel A. 1894b. Quelques fourmis de Madagascar (récoltées par M. le Dr. Voltzkow); de Nouvelle Zélande (récoltées par M. W. W. Smith); de Nouvelle Calédonie (récoltées par M. Sommer); de Queensland (Australie) (récoltées par M. Wiederkehr); et de Perth (Australie occidentale) (récoltées par M. Chase). Annales de la Société Entomologique de Belgique 38: 226–237.
  • Forel A. 1895a. Nouvelles fourmis de diverses provenances, surtout d'Australie. Annales de la Société Entomologique de Belgique 39: 41–49.
  • Forel A. 1895b. Südpalaearctische Ameisen. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 9: 227–234.
  • Forel A. 1902. Myrmicinae nouveaux de l'Inde et de Ceylan. Revue Suisse de Zoologie 10: 165–249.
  • Forel A. 1904a. Hymenoptera, Formicidae. Pp. 13–15, In: Wasmann E. 1904. Termitophilen aus dem Sudan, In: Jägerskiöld L.A. 1904. Results of the Swedish Zoological Expedition to Egypt and the White Nile, 1901. Part 1 (no. 13). Uppsala: Library of the Royal University of Uppsala, 21 pp.
  • Forel A. 1904b. Dimorphisme du mâle chez les fourmis et quelques autres notices myrmécologiques. Annales de la Société Entomologique de Belgique 48: 421–425.
  • Forel A. 1904c. Miscellanea myrmécologiques. Revue Suisse de Zoologie 12: 1–52.
  • Forel A. 1905. Miscellanea myrmécologiques II. Annales de la Société Entomologique de Belgique 49: 155–185.
  • Forel A. 1907a. Formicides du Musée National Hongrois. Annales Historico-Naturales Musei Nationalis Hungarici 5: 1–42.
  • Forel A. 1907b. Fourmis nouvelles de Kairouan et d'Orient. Annales de la Société Entomologique de Belgique 51: 201–208.
  • Forel A. 1910a. Glanures myrmécologiques. IV. Fourmis de Crète. Annales de la Société Entomologique de Belgique 54: 21–23.
  • Forel A. 1910b. Glanures myrmécologiques. V. Notices sur les fourmis de Grèce. Annales de la Société Entomologique de Belgique 54: 23–27.
  • Forel A. 1910c. Glanures myrmécologiques. Annales de la Société Entomologique de Belgique 54: 6–32.
  • Forel A. 1910d. Note sur quelques fourmis d'Afrique. Annales de la Société Entomologique de Belgique 54: 421–458.
  • Forel A. 1911a. Fourmis nouvelles ou intéressantes. Bulletin de la Société Vaudoise des Sciences Naturelles 47: 331–400.
  • Forel A. 1911b. Die Ameisen des K. Zoologischen Museums in München. Sitzungsberichte der Mathematischen-Physikalischen Klasse der Königlich Bayerischen Akademie der Wissenschaften zu München 11: 249–303.
  • Forel A. 1912. Formicides néotropiques. Part V. 4me sous-famille Dolichoderinae Forel. Mémoires de la Société Entomologique de Belgique 20: 33–58.
  • Forel A. 1913a. Fourmis de la faune méditerranéenne récoltées par MM. U. et J. Sahlberg. Revue Suisse de Zoologie 21: 427–438.
  • Forel A. 1913b. Fourmis d'Argentine, du Brésil, du Guatémala & de Cuba reçues de M. M. Bruch, Prof. v. Ihering, Mlle Baez, M. Peper et M. Rovereto. Bulletin de la Société Vaudoise des Sciences Naturelles 49: 203–250.
  • Forel A. 1914. Formicides d'Afrique et d'Amérique nouveaux ou peu connus. Bulletin de la Société Vaudoise des Sciences Naturelles 50: 211–288.
  • Ghahari H., Collingwood C. 2013. A study on the ants (Hymenoptera: Vespoidea: Formicidae) from western Iran. Acta Phytopathologica et Entomologica Hungarica 48(1): 155–164.
  • Grove A.T., Rackham O. 1993. Threatened landscapes in the Mediterranean: examples from Crete. Landscape and Urban Planning 24: 279–292.
  • Guérin-Méneville F.E. 1852. Notice sur une nouvelle espèce de fourmi découverte à Saint-Domingue par M. Auguste Sallé, et qui fait son nid dans les plaines marécageuses, sur les buissons. Revue et Magasin de Zoologie Pure et Appliquée 4(2): 73–79.
  • Henin J.M., Paiva M.R., Collingwood C.A. 2001. The Iberian Leptothorax Mayr 1855 (Hymenoptera, Formicidae): discovery of a new species and first record of L. Atlantis Santschi 1911. Boletim da Sociedade Portuguesa de Entomologia 196: 161–166.
  • Higgins M.D. 2009. Greek Is., Geology. In: Gillespie R.G., Clague D.A. (Ed.), Encyclopedia of Isl. Berkeley: University of California Press: 392–396.
  • Hijmans R.J., Guarino L., Mathur P., Jarvis A., Rojas E. 2011. DIVA-GIS. Version 7.5. 0.0.
  • Ionescu–Hirsch A. 2009. An annotated list of Camponotus of Israel (Hymenoptera: Formicidae), with a key and descriptions of new species. Israel Journal of Entomology 39: 57–98.
  • Jerdon T.C. 1851. A catalogue of the species of ants found in Southern India. Madras Journal of Literature and Science 17: 103–127.
  • Jucker C., Rigato F., Regalin R. 2008. Exotic ant records from Italy (Hymenoptera, Formicidae). Bollettino di Zoologia Agraria e di Bachicoltura 40: 99–107.
  • Kaltsas D., Trichas A., Kougioumoutzis K., Chatzaki M. 2013. Ground beetles respond to grazing at assemblage level, rather than species-specifically: the case of Cretan shrublands. Journal of Insect Conservation 14: 681–697.
  • Karaman M.G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8: 151–162.
  • Karaman M.G. 2011. Zoogeography, diversity and altitudinal distribution of ants (Hymenoptera: Formicidae) in the Mediterranean and the oro-Mediterranean parts of Montenegro. North-Western Journal of Zoology 7: 26–34.
  • Karaman C., Aktaç N. 2013. Descriptions of four new species of Camponotus Mayr (Hymenoptera: Formicidae), with a key for the worker caste of the Camponotus of Turkey. Journal of the Kansas Entomological Society 86(1): 36–56.
  • Karaman C., Aktaç N., Kiran K. 2011. Ants of the genus Camponotus Mayr 1861 (Hymenoptera: Formicidae) in the Kaz Mountains, Turkey, with descriptions of sexuals of Camponotus candiotes Emery, 1894 and Camponotus ionius Emery, 1920. Turkish Journal of Zoology 35(2): 183–197.
  • Karavaiev V. 1912. Ameisen aus dem paläarktischen Faunengebiete. Russkoe entomologischeskoe Obozrenie 12: 581–596.
  • Karavaiev V. 1926. Beiträge zur Ameisenfauna des Kaukasus, nebst einigen Bemerkungen über andere palaearktische Formen. (Schluss). Konowia 5: 187–199.
  • Karavaiev V. 1927a. Myrmekologische Miszellen. Izvestiia Akademii nauk SSSR. Seriia biologicheskaia 27: 104–112.
  • Karavaiev V. 1927b. Ameisen aus dem paläarktischen Gebiet. II. Zbirnyk Prats' Zoolohichnoho Muzeyu 2: 89–104.
  • Karavaiev V. 1931 (1930). Myrmekologische Fragmente, III. Zoologischer Anzeiger 92: 309–317.
  • Karavaiev V. 1932. Zwei neue Ameisen aus Aserbeidschan (Transkaukasien). Zoologischer Anzeiger 98: 248–250.
  • Kiran K., Karaman C. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548(1): 1–38.
  • Kratochvíl J., Novák V., Snoflák J. 1944. Mohelno. Soubor práci venoványch studiu vyznamne památky prírodní. 5. Hymenoptera - Aculeata. Formicidae - Apidae - Vespidae. Archiv Svazu na Ochranu Prírody a Domoviny na Moravé 6: 1–155.
  • Krausse A.H. 1910. Über Stridulationstöne bei Ameisen. Zoologischer Anzeiger 35: 523–526.
  • Krausse A.H. 1911 ("1909"). Über Messor structor Ltr. und einige andere Ameisen auf Sardinien. Bullettino della Società Entomologica Italiana 41: 14–18.
  • Krausse A.H. 1912. Eine neue Ameisenform von Sardinien (Pheidole pallidula v. n. Emeryi m.). Internationale Entomologische Zeitschrift 6: 169.
  • Krijgsman W., Hilgen F.J., Raffi I., Sierro F.J., Wilson D.S. 1999. Chronology, causes and progression of the Messinian salinity crisis. Nature 400: 652–655.
  • Kutter H. 1950. Über zwei neue Ameisen. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 23: 337–346.
  • Kutter H. 1973. Zur Taxonomie der Gattung Chalepoxenus (Hymenoptera, Formicidae, Myrmicinae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 46: 269–280.
  • Kuznetsov-Ugamsky N.N. 1923. The ant fauna of Tashkent district. [In Russian]. Trudy Turkestanskogo Nauchnogo Obshchestva 1: 239–558.
  • Kuznetsov-Ugamsky N.N. 1929. Die Ameisenfauna Daghestans. Zoologischer Anzeiger 83: 34–45.
  • Lapeva-Gjonova A., Antonova V., Radchenko A.G., Atanasova M. 2010. Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria. ZooKeys 62: 1–124.
  • Latreille P.A. 1798. Essai sur l'histoire des fourmis de la France. Brive: F. Bourdeaux, 50 pp.
  • Latreille P.A. 1802. Histoire naturelle des fourmis, et recueil de mémoires et d'observations sur les abeilles, les araignées, les faucheurs, et autres insectes. Paris: Impr. Crapelet (chez T. Barrois), xvi + 445 pp.
  • Leach W.E. 1825. Descriptions of thirteen species of Formica and three species of Culex found in the environs of Nice. Zoological Journal. London 2: 289–293.
  • Legakis A. 1983. Contribution to the zoogeography of ants (Hymenoptera, Formicidae) in the Greek Isl. Biologia gallo-hellenica 10: 253–257.
  • Legakis A. 2011. Annotated list of the ants (Hymenoptera, Formicidae) of Greece. Hellenic Zoological Archives 7: 1–55.
  • Legakis A., Kollaros D., Kypriotakis Z., Paragamian K., Trichas A., Voreadou C. 1993 Ecological assessment of the coasts of Crete (Greece). Coastal Management 21 143–154
  • Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Holmiae: L. Salvii, 824 pp.
  • Linnaeus C. 1767. Systema naturae, per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. Tomus I. Pars 2. Editio duodecima, reformata. Holmiae: L. Salvii: 533–1327.
  • Lomnicki J. 1925. Plagiolepis vindobonensis n. sp. (Hym. Formicidae). Polskie Pismo Entomologiczne 4: 77–79.
  • Losana M. 1834. Saggio sopra le formiche indigene del Piemonte. Memorie della Reale Accademia delle Scienze di Torino 37: 307–333.
  • Lucas H. 1849. Exploration scientifique de l'Algérie pendant les années 1840, 1841, 1842. Sciences physiques. Zoologie. III. Histoire naturelle des animaux articulés. Troisième partie - Insectes (Suite). Paris: A. Bertrand, 527 pp.
  • Lucas M.H. 1854. Essai sur les animaux articulés qui hebitent l'ile de Crète. Revue de Magasin de Zoologie 6: 487–491.
  • Mayr G. 1853a. Einige neue Ameisen. Verhandlungen der Zoologisch-Botanischen Vereins in Wien 2: 143–150.
  • Mayr G. 1853b. Beschreibungen einiger neuer Ameisen. Verhandlungen der Zoologisch-Botanischen Vereins in Wien 3: 277–286.
  • Mayr G. 1855. Formicina austriaca. Beschreibung der bisher im österreichischen Kaiserstaate aufgefundenen Ameisen, nebst Hinzufügung jener in Deutschland, in der Schweiz und in Italien vorkommenden Arten. Verhandlungen der Zoologisch-Botanischen Vereins in Wien 5: 273–478.
  • Mayr G. 1861. Die europäischen Formiciden. Nach der analytischen Methode bearbeitet. Wien: C. Gerolds Sohn, 80 pp
  • Mayr G. 1868. Formicidae novae Americanae collectae a Prof. P. de Strobel. Annuario della Società dei Naturalisti e Matematici, Modena 3(1): 61–178.
  • Mayr G. 1870. Neue Formiciden. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 20: 939–996.
  • Mayr G. 1879. Beiträge zur Ameisen-Fauna Asiens. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 28: 645–686.
  • Mayr G. 1904. Formiciden aus Ägypten und dem Sudan, In: Jägerskiöld L.A. 1904. Results of the Swedish Zoological Expedition to Egypt and the White Nile, 1901. Part 1 (no. 9). Uppsala: Library of the Royal University of Uppsala, 11 pp.
  • Menozzi C. 1921. Formiche dei dintorni di Sambiase di Calabria. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura 15: 24–32.
  • Menozzi C. 1923 ("1922"). Nota su un nuovo genere e nuova specie di formica parassita. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 61: 256–260.
  • Menozzi C. 1929. Ricerche faunistiche nelle isole italiane dell'Egeo. Imenotteri (Formiche). Archivo Zoologico Itaiano 13(1–2): 145–146.
  • Menozzi C. 1933. Le formiche della Palestina. Memorie della Società Entomologica Italiana 12: 49–113.
  • Menozzi C. 1936. Nuovi contributi alla conoscenza della fauna delle Isole italiane dell'Egeo. VI. Hymenoptera - Formicidae. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura 29: 262–311.
  • Motschoulsky V.T. 1839. Insectes du Caucase et des provinces transcaucasiennes. Bulletin de la Société Impériale des Naturalistes de Moscou 12: 44–68.
  • Müller G. 1923. Le formiche della Venezia Giulia e della Dalmazia. Bollettino della Società adriatica di scienze naturali in Trieste 28(1): 11–180.
  • Neuenschwander P., Bigler F., Delucchi V., Michelakis S. 1983. Natural enemies of preimaginal stages of Dacus oleae Gmel. (Dipt. Tephritidae) in Western Crete. I. Bionomics and phenologies. Bollettino del Laboratorio di Zoologia Generale e Agraria 40: 3–32.
  • Novák V., Sadil J. 1941. Klíc k urcování mravencu strední Evropy se zvlástním zretelem k mravencí zvírene Cech a Moravy. Entomologické Listy 4: 65–115.
  • Nylander W. 1846a. Adnotationes in monographiam formicarum borealium Europae. Acta Societatis Scientiarum Fennicae 2: 875–944.
  • Nylander W. 1846b. Additamentum adnotationum in monographiam formicarum borealium Europae. Acta Societatis Scientiarum Fennicae 2: 1041–1062
  • Nylander W. 1849 ("1848"). Additamentum alterum adnotationum in monographiam formicarum borealium. Acta Societatis Scientiarum Fennicae 3: 25–48.
  • Nylander W. 1856. Synopsis des Formicides de France et d'Algérie. Annales des Sciences Naturelles, Zoologie 5(4): 51–109.
  • Olivier A.G. 1792. Encyclopédie méthodique. Histoire naturelle. Insectes. Tome 6. (pt. 2). Paris: Panckoucke: 369–704.
  • Özdikmen H. 2010. New names for some preoccupied specific and subspecific epithets in the family Formicidae (Hymenoptera). Munis Entomology and Zoology 5(supplement): 986–1000.
  • Passalacqua N.G. 2015. On the definition of element, chorotype and component in biogeography. Journal of Biogeography 42 (4): 611–618.
  • Perris E. 1878 (1877). Rectifications et additions à mes promenades entomologiques. Annales de la Société Entomologique de France 7(5): 379–386.
  • Perissoratis C., Conispoliatis N. 2003. The impacts of sea-level changes during latest Pleistocene and Holocene times on the morphology of the Ionian and Aegean seas (SE Alpine Europe). Marine Geology 196: 145–156.
  • Petrov I. 2006. The ants of Serbia and Montenegro. Posebna Izdanja. Srpska Akademija Nauka i Umetnosti 656: 1–136.
  • Poulakakis N., Kapli P., Lymberakis P., Trichas A., Vardinoyiannis K., Sfenthourakis S., Mylonas M. 2015. A review of phylogeographic analyses of animal taxa from the Aegean and surrounding regions. Journal of Zoological Systematics and Evolutionary Research 53(1): 18–32.
  • Rackham O., Moody J.A. 1996. The Making of the Cretan Landscape. Manchester University Press: Manchester.
  • Radchenko A.G. 1992. Ants of the genus Tetramorium (Hymenoptera, Formicidae) of the USSR fauna. Report 2. [In Russian.]. Zoologicheskii Zhurnal 71(8): 50–58.
  • Radchenko A.G. 1996a Ants of the genus Plagiolepis Mayr (Hymenoptera, Formicidae) of the central and southern Palearctic [In Russian]. Entomologicheskoe Obozrenie 75:178–187.
  • Radchenko A.G. 1996b. A key to the ant genus Camponotus (Hymenoptera, Formicidae) in Palearctic Asia. [In Russian]. Zoologicheskii Zhurnal 75: 1195–1203.
  • Radchenko A.G. 1997. Review of the ants of scabriceps group of the genus Monomorium Mayr (Hymenoptera, Formicidae). Annales Zoologici 46(3–4): 211–224.
  • Radchenko A.G., Arakelian G.R. 1990. Ants of the group Tetramorium ferox Ruzsky (Hymenoptera, Formicidae) from Crimea and the Caucasus. [In Russian]. Biologicheskii Zhurnal Armenii 43: 371–378.
  • Raulin V. 1861. Description physique de l'île de Crète. Partie zoologique. Actes de la Société linnéenne de Bordeaux 24: 643–708.
  • Roger J. 1859. Beiträge zur Kenntniss der Ameisenfauna der Mittelmeerländer. I. Berliner Entomologische Zeitschrift 3: 225–259.
  • Roger J. 1860. Die Ponera-artigen Ameisen. Berliner Entomologische Zeitschrift 4: 278–312.
  • Roger J. 1862. Beiträge zur Kenntniss der Ameisenfauna der Mittelmeerländer. II. Berliner Entomologische Zeitschrift 6: 255–262.
  • Roger J. 1863. Die neu aufgeführten Gattungen und Arten meines Formiciden-Verzeichnisses nebst Ergänzung einiger früher gegebenen Beschreibungen. Berliner Entomologische Zeitschrift 7: 131–214.
  • Ruzsky M. 1896. Verzeichniss der Ameisen des östlichen Russlands und des Uralgebirges. Berliner Entomologische Zeitschrift 41: 67–74.
  • Ruzsky M. 1902. Material on the ant fauna of the Caucasus and the Crimea. [In Russian]. Protokoly Obshchestva Estestvoispytatelei pri Imperatorskom Kazanskom Universitete 206(suppl.): 1–33.
  • Ruzsky M. 1903. Essay on the myrmecofauna of the Kirghiz steppe. [In Russian]. Trudy Russkago Entomologicheskago Obshchestva 36: 294–316.
  • Ruzsky M. 1905. The ants of Russia. (Formicariae Imperii Rossici). Systematics, geography and data on the biology of Russian ants. Part I. [In Russian]. Trudy Obshchestva Estestvoispytatelei pri Imperatorskom Kazanskom Universitete 38(4-6): 1–800.
  • Ruzsky M. 1914. Eine neue Ameisenform aus dem europäischen Russland. Russkoe Entomologicheskoe Obozrenie 14: 323.
  • Ruzsky M. 1915. Material on Siberian myrmecology. First output. On the myrmecofauna of Tomsk province and certain other Siberian localities. (From research in 1914-1915). [In Russian]. Izvestiya Imperatorskago Tomskago Universiteta 64(5): 1–14.
  • Salata S., Borowiec L. 2015a. A taxonomic revision of the genus Oxyopomyrmex André, 1881 (Hymenoptera: Formicidae). Zootaxa (Monograph) 4025(1): 1–66.
  • Salata S., Borowiec L. 2015b. Redescription of Temnothorax antigoni (Forel, 1911) and description of its new social parasite Temnothorax curtisetosus sp. n. from Turkey (Hymenoptera, Formicidae). ZooKeys 523: 129–148.
  • Salata S., Borowiec L. 2015c. Redescription of Crematogaster cypria Santschi, 1930, new status, with description of two new related species from Greece and Turkey (Hymenoptera, Formicidae). ZooKeys 505: 59–78.
  • Salata S., Borowiec L. 2016. A new species of the Aphaenogaster cecconii group (Hymenoptera: Formicidae) from Rhodes. Zootaxa 4170(1): 194–200.
  • Salata S., Borowiec L. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici 67(2): 279–313.
  • Salata S., Borowiec L. 2019. Preliminary contributions toward a revision of Greek Messor Forel, 1890 (Hymenoptera: Formicidae). Turkish Journal of Zoology 43: 52–67.
  • Salata S., Borowiec L. 2018a. A new species of the ant genus Lasius Fabricius, 1804 from Crete (Hymenoptera, Formicidae). ZooKeys 789: 139–159.
  • Salata S., Borowiec L. 2018b. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom, Entomology, 27(008): 1–51.
  • Salata S., Georgiadis C., Borowiec L. 2018. Invasive ant species (Hymenoptera: Formicidae) of Greece and Cyprus. North-Western Journal of Zoology: e171204.
  • Salata S., Borowiec L., Radchenko A.G. 2018a. Description of Plagiolepis perperamus, a new species from East-Mediterranean and redescription of Plagiolepis pallescens Forel, 1889 (Hymenoptera: Formicidae). Annales Zoologici 68(4): 809–824.
  • Salata S., Borowiec L., Trichas A. 2018b. Taxonomic revision of the Cretan fauna of the genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae), with notes on the endemism of ant fauna of Crete. Annales Zoologici 68(4): 769–808.
  • Santschi F. 1908. Nouvelles fourmis de l'Afrique du Nord (Égypte, Canaries, Tunisie). Annales de la Société Entomologique de France 77: 517–534.
  • Santschi F. 1909. Leptothorax rottenbergi et espèces voisines. Revue Suisse de Zoologie 17: 459–482.
  • Santschi F. 1911 (1910). Nouvelle fourmis d'Afrique. Annales de la Société Entomologique de France 79: 351–369.
  • Santschi F. 1912. Quelques nouvelles variétés de fourmis africaines. Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 3: 147–149.
  • Santschi F. 1913. Glanures de fourmis africaines. Annales de la Société Entomologique de Belgique 57: 302–314.
  • Santschi F. 1914. Formicides de l'Afrique occidentale et australe du voyage de Mr. le Professeur F. Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura, Portici 8: 309–385.
  • Santschi F. 1915. Nouvelles fourmis d'Afrique. Annales de la Société Entomologique de France 84: 244–282.
  • Santschi F. 1916. Formicides sudaméricains nouveaux ou peu connus. Physis 2: 365–399.
  • Santschi F. 1917a. Acantholepi Frauenfeldi Mayr et ses variétés. Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 8: 42–48.
  • Santschi F. 1917b. Races et variétés nouvelles du Messor barbarus L. Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 8: 89–94.
  • Santschi F. 1919a. Nouveaux formicides de la République Argentine. Anales de la Sociedad Cientifica Argentina 87: 37–57.
  • Santschi F. 1919b. Trois nouvelles fourmis des Canaries. Boletín de la Real Sociedad Española de Historia Natural 19: 405–407.
  • Santschi F. 1919c. Nouveaux genre et sous-genre de fourmis barbaresques (Hym.). Bulletin de la Société Entomologique de France 1919: 90–92.
  • Santschi F. 1920. Cinq nouvelles notes sur les fourmis. Bulletin de la Société Vaudoise des Sciences Naturelles 53: 163–186.
  • Santschi F. 1921a. Notes sur les fourmis paléarctiques. II. Fourmis d'Asie Mineure récoltées par M. H. Gadeau de Kerville. Boletín de la Real Sociedad Española de Historia Natural 21: 110–116.
  • Santschi F. 1921b. Nouvelles fourmis paléarctiques. 3ème. note. Boletín de la Real Sociedad Española de Historia Natural 21: 165–170.
  • Santschi F. 1923a. Notes sur les fourmis paléarctiques. 4ème note. Boletín de la Real Sociedad Española de Historia Natural 23: 133–137.
  • Santschi F. 1923b. Pheidole et quelques autres fourmis néotropiques. Annales de la Société Entomologique de Belgique 63: 45–69.
  • Santschi F. 1923c. Messor et autres fourmis paléarctiques. Revue Suisse de Zoologie 30: 317–336.
  • Santschi F. 1925. Fourmis d'Espagne et autres espèces paléarctiques (Hymenopt.). EOS. Revista Española de Entomología 1: 339–360.
  • Santschi F. 1926a.– Travaux scientifiques de l'Armée d'Orient (1916–1918). Fourmis. Bulletin du Muséum national d'histoire naturelle 1926(5): 286–293.
  • Santschi F. 1926b. Description de nouveaux Formicides éthiopiens (IIIme partie). Revue Zoologique Africaine 13: 207–267.
  • Santschi F. 1927a. Revision des Messor du groupe instabilis Sm. (Hymenopt.). Boletín de la Real Sociedad Española de Historia Natural 27: 225–250.
  • Santschi F. 1927b. Révision myrmécologique. Bulletin et Annales de la Société Entomologique de Belgique 67: 240–248.
  • Santschi F. 1927c. Notes myrmécologiques. Bulletin de la Société Entomologique de France 1927: 126–128.
  • Santschi F. 1927d. A propos du Tetramorium caespitum L. Folia Myrmecologica et Termitologica 1: 52–58.
  • Santschi F. 1928. Nouvelles fourmis de Chine et du Turkestan Russe. Bulletin et Annales de la Société Entomologique de Belgique 68: 31–46.
  • Santschi F. 1929a. Étude sur les Cataglyphis. Revue Suisse de Zoologie 36: 25–70.
  • Santschi F. 1929b. Nouvelles fourmis de la République Argentine et du Brésil. Anales de la Sociedad Cientifica Argentina 107: 273–316.
  • Santschi F. 1929c. Fourmis du Maroc, d'Algérie et de Tunisie. Bulletin et Annales de la Société Entomologique de Belgique 69: 138–165.
  • Santschi F. 1931. Inventa entomologica itineris Hispanici et Maroccani, quod a. 1926 fecerunt Harald et Håkan Lindberg. Fourmis du Bassin Méditerranéen occidental et du Maroc récoltées par MM. Lindberg. Commentationes Biologicae 3(14): 1–13.
  • Santschi F. 1934. Fourmis d'une croisière. Bulletin et Annales de la Société Entomologique de Belgique 74: 273–282.
  • Santschi F. 1936. Étude sur les fourmis du genre Monomorium Mayr. Bulletin de la Société des Sciences Naturelles du Maroc 16: 32–64.
  • Santschi F. 1939. Notes sur des Camponotus et autres fourmis de l'Afrique Mineure. Bulletin de la Société des Sciences Naturelles du Maroc 19: 66–87.
  • Sarnat E.M., Fischer G., Guénard B., Economo E.P. 2015. Introduced Pheidole of the world: taxonomy, biology and distribution. ZooKeys 543: 1–109.
  • Schmidt C. 2013. Molecular phylogenetics of ponerine ants (Hymenoptera: Formicidae: Ponerinae). Zootaxa 3647(2): 201–250.
  • Schule W. 1993. Mammals, vegetation and the initial human settlement of the Mediterranean Islands – a paleoecological approach. Journal of Biogeography 20: 399–412.
  • Schulz A. 1996. Tetramorium rhenanum nov. spec. vom "Mittleren Rheintal" in Deutschland (Hymenoptera: Formicidae). Linzer Biologische Beiträge 28: 391–412.
  • Scopoli J.A. 1763. Entomologia carniolica exhibens insecta Carnioliae indigena et distributa in ordines, genera, species, varietates. Methodo Linnaeana. Vindobonae: J. Trattner, xxxvi + 420 pp.
  • Seifert B. 1992. A taxonomic revision of the Palaearctic members of the ant subgenus Lasius s. str. (Hymenoptera, Formicidae). Abhandlungen und Berichte des naturkundemuseums Görlitz 66: 1–67
  • Seifert B. 2003. The ant genus Cardiocondyla (Insecta: Hymenoptera: Formicidae) – a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambouloffii, C. wroughtonii, C. emeryi and C. minutior species groups. Annalen des Naturhistorischen Museums in Wien 104B: 203–338.
  • Seifert B. 2016. Inconvenient hyperdiversity – the traditional concept of "Pheidole pallidula" includes four cryptic species (Hymenoptera: Formicidae). Soil Organisms 88(1): 1–17.
  • Seifert B., Galkowski C. 2016. The Westpalaearctic Lasius paralienus complex (Hymenoptera: Formicidae) contains three species. Zootaxa 4132(1): 44–58.
  • Shuckard W.E. 1838. Description of a new species of Myrmica which has been found in houses both in the Metropolis and Provinces. Magazine of Natural History (2)2: 626–627
  • Smith F. 1857. Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace. Journal and Proceedings of the Linnean Society of London. Zoology 2: 42–88.
  • Smith F. 1858. Catalogue of hymenopterous insects in the collection of the British Museum. Part VI. Formicidae. London: British Museum, 216 pp.
  • Smith F. 1860. Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the Isl of Bachian, Kaisaa, Amboyna, Gilolo, and at Dory in New Guinea. Journal and Proceedings of the Linnean Society of London. Zoology 5(17b) (suppl. to vol. 4): 93–143.
  • Smith F. 1861. Descriptions of some new species of ants from the Holy Land, with a synonymic list of others previously described. Journal and Proceedings of the Linnean Society of London. Zoology 6: 31–35.
  • Smith F. 1862. Descriptions of new species of aculeate Hymenoptera, collected at Panama by R. W. Stretch, Esq., with a list of described species, and the various localities where they have previously occurred. Transactions of the Entomological Society of London 1(3): 29–44.
  • Snelling R.R. 1974. Studies on California ants. 8. A new species of Cardiocondyla (Hymenoptera: Formicidae). Journal of the New York Entomological Society 82: 76–81.
  • Soudek S. 1925. Four new European ants. Entomologist's Record and Journal of Variation 37: 33–37.
  • Spinola M. 1808. Insectorum Liguriae species novae aut rariores, quae in agro ligustico nuper detexit, descripsit et iconibus illustravit Maximilianus Spinola, adjecto catalogo specierum auctoribus jam enumeratarum, quae in eadam regione passim occurrent. Tom. II. Fasc. 4. Genova: Y. Gravier: 207–262.
  • Steiner F.M., Csősz S., Markó B., Gamisch A., Rinnhofer L., Folterbauer C., Hammerle S., Stauffer C., Arthofer W., Schlick-Steiner B.C. 2018. Turning one into five: integrative taxonomy uncovers complex evolution of cryptic species in the harvester ant Messor "structor". Molecular Phylogenetics and Evolution 127: 387–404.
  • Stitz H. 1917. Ameisen aus dem westlichen Mittelmeergebiet und von den Kanarischen Inseln. Mitteilungen aus dem Zoologischen Museum in Berlin 8: 333–353.
  • Stitz H. 1928. Zoologische Streifzüge in Attika, Morea und besonders auf der Insel Crete. VIII. Hymenoptera: Formicidae. Naturwissenschaftlicher Verein zu Bremen 27: 90–91.
  • Stitz H. 1930. XXXI. Fam. Formicidae. Ameisen. Pp. 521-563 in: Schmiedeknect O. (Ed.) 1930. Die Hymenopteren Nord- und Mitteleuropas. Zweite Auflage. Jena: G. Fischer, x + 1062 pp.
  • Stitz H. 1939. Die Tierwelt Deutschlands und der angrenzenden Meersteile nach ihren Merkmalen und nach ihrer Lebensweise. 37. Theil. Hautflüger oder Hymenoptera. I: Ameisen oder Formicidae. Jena: G. Fischer, 428 pp.
  • Tăuşan I., Markó B. 2011. New records of the pharaoh ant Monomorium pharaonis (Linnaeus, 1758) (Hymenoptera: Formicidae) in Romania. Entomologica Romanica 16: 23–26.
  • Tohmé G., Tohmé H. 1981. Les fourmis du genre Messor en Syrie. Position systématique. Description de quelques ailés et de formes nouvelles. Répartition géographique. Ecologia Mediterranea 7(1): 139–153.
  • Triantis K.A., Mylonas M. 2009. Greek Is., Biology. In: Gillespie R.G., Clague D.A. (Eds.), Encyclopedia of Isl. Berkeley: University of California Press: 388–392.
  • Vesnić A. 2011. Revidirani sistematski prijegled mrava Bosne i Hercegovine. Unutar: S. Lelo (urednik), Fauna Bosne i Hercegovine – Biosistematski pregledi. 7. izmijenjeno i popravljeno interno izdanje Udruženja za inventarizaciju i zaštitu životinja, Ilijaš, Kanton Sarajevo: 205–207.
  • Vigna Taglianti A., Audisio P.A., Biondi M., Bologna M.A., Carpaneto G.M., De Biase A., Fattorini S., Piatella E., Sindaco R., Venchi A., Zapparoli M. 1999. A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palearctic region. Biogeographia 20: 31–59.
  • Vogiatzakis I.N., Pungetti G. Mannion A.M. 2008. Mediterranean island landscapes: natural and cultural approaches. In: Landscape Series, vol. 9. Springer Publishing.
  • Wagner H.C., Arthofer W., Seifert B., Muster C., Steiner F.M., Schlick-Steiner B.C. 2017. Light at the end of the tunnel: integrative taxonomy delimits cryptic species in the Tetramorium caespitum complex (Hymenoptera: Formicidae). Myrmecological News 25: 95–129.
  • Warchałowski A. 1976. Biogeographische Studien über die Blattkäfer der Pontischen Provinz (Coleoptera, Chrysomelidae). Polskie Pismo Entomologiczne 46: 29–94.
  • Ward P.S. (Ed.) 2013. AntWeb. Available from: https://www.antweb.org/. Accesed 21 II 2017.
  • Ward P.S. 2014. The phylogeny and evolution of ants. Annual Review of Ecology, Evolution, and Systematics 45: 23–43.
  • Ward P.S., Brady S.G., Fisher B.L., Schultz T.R. 2010. Phylogeny and biogeography of dolichoderine ants: effects of data partitioning and relict taxa on historical inference. Systematic Biology: syq012.
  • Ward P.S., Brady S.G., Fisher B.L., Schultz T.R. 2015. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology 40(1): 61–81.
  • Ward P.S., Blaimer B.B., Fisher B.L. 2016. A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the genera Colobopsis and Dinomyrmex. Zootaxa 4072(3): 343–357.
  • Wasmann E. 1909. Über den Ursprung des sozialen Parasitismus, der Sklaverei und der Myrmekophilie bei den Ameisen. Biologisches Centralblatt 29: 587–604.
  • Wetterer J.K., Wild A.L., Suarez A.V., Roura–Pascual N., Espadaler X. 2009. Worldwide spread of the Argentine ant, Linepithema humile (Hymenoptera: Formicidae). Myrmecological News 12: 187–194.
  • Wetterer J.K. 2010. Worldwide spread of the pharaoh ant Monomorium pharaonis (Hymenoptera: Formicidae). Myrmecological News 13: 115–129.
  • Wheeler W.M. 1903. Some new gynandromorphous ants, with a review of the previously recorded cases. Bulletin of the American Museum of Natural History 19: 653–683.
  • Wheeler WM. 1910. Ants: their structure, development and behavior. New York: Columbia University Press, xxv + 663 ss.
  • Wheeler W.M. 1921. Chinese ants. Bulletin of the Museum of Comparative Zoology 64: 529–547.
  • Wheeler W.M. 1922. Ants of the American Museum Congo expedition. New York. Bulletin of the American Museum of Natural History 1139.
  • Wheeler W.M. 1926. Ants of the Balearic Isl. Folia Myrmecologica et Termitologica 1: 1–6.
  • Zimmermann S. 1935 ("1934"). Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 84: 1–65.