2. Baetis (Tenuibaetis) hissaricus Novikova 1991

(Figs 44–59)

Baetis ursinus hissaricus Novikova 1991: 793 (larva, ♁ imago, ♀ imago, subimago); Kluge 1995: 12 (type specimens).

Baetis (Baetis) ursinus hissaricus: Kluge 1997b: 194 (larva).

Baetis (Tenuibaetis) hissaricus: Kluge 2022: 166 (subimago).

Material examined (ZIN, holotype and paratypes). TAJIKISTAN: Garm, small stream, 15. VIII.1981, coll. N. Kluge: 18 larvae; basin of river Sardai-Miona 15 km upstream Ramit, 15–22. VIII.1981, coll. N. Kluge & E. Novikova: 1 L-S♁, 3 L-S ♀, 15 larvae; valley Divdara, stream Zangob, 12 km upstream Ramit by river Sardai-Miona, 18–21. VIII.1981, coll. N. Kluge & E. Novikova: 2 L-S-I♁, 5 L-S-I ♀, 3 L-S♁, 10 L-S ♀.

Additional descriptions

Larva. CUTICULAR COLORATION (Figs 44, 47–49: Described and figured by Novikova (1991: figs 1–16); pronotum with darker sigilla on lighter background (Novikova 1991: figs 1–2).

HYPODERMAL COLORATION: Not expressed.

SHAPE AND SETATION: Mouthparts— Figs 44–46. Hind protoptera present (as in Fig. 2).

Two-channel setae forming row along outer side of femur (including two subapical setae) elongate, blunt, distally widened and flattened; stout setae on anterior side of femur two-channel, from moderately long to short (Figs 47–49).

Tibia of each leg with small, stout two-channel setae on inner side and along patella-tibial suture (Fig. 51). Stout setae on outer side of tibia pseudo-bifurcate and very dense, forming not one row, but stripe of irregularly situated setae 2–4 setae width (Fig. 50).

Denticles on posterior margins of terga I–X bluntly triangular (Novikova 1991: fig. 30). Denticles on posterior margin of sternum small and irregular, present only on sternum IX, sometimes also on sterna VIII and VII; in male row of denticles not interrupted by protogonostyli. Paraproct with denticles on most part of inner margin (as in Fig. 30). Paracercus significantly smaller than cerci.

Subimago. CUTICULAR COLORATION: Head colorless, antennae brown. Pronotum light brown. Mesonotum with medioscutum and submedioscutum mostly ochre, lateral areas mostly light brownish (Fig. 59). Thoracic pleura and sterna with light brownish and dark brown areas (as in Fig. 215). Legs of all pairs nearly colorless, with small, contrasting, brown marking near base of tibia (Fig. 58). All abdominal terga and sterna I–X light brownish.

HYPODERMAL COLORATION: As in imago.

Imago, male. Described by Novikova (1991). Middle and hind tarsi with 2 apical spines (on 1st+2nd and 3rd segments). Gonovectes gently bent, brown; sterno-styligeral muscle vestigial (Fig. 54).

Imago, female. Described by Novikova (1991). All tarsi with 2 apical spines (fore tarsus on 2nd and 3rd segments, middle and hind tarsi on 1st+2nd and 3rd segments).

Dimension. Fore wing length 4–5 mm.

Distribution. Mountains of Central Asia.

Comparison. Larva of B. (T.) hissaricus differs from other examined species by very dense stripe of pseudo-bifurcate setae on outer sides of tibiae (Figs 50–51).