Published December 15, 2016 | Version v1

Otospermophilus beecheyi (Rodentia: Sciuridae)

Description

Smith, Jennifer E., Long, Douglas J., Russell, Imani D., Newcomb, Kate Lee, Muñoz, Valeska D. (2016): Otospermophilus beecheyi (Rodentia: Sciuridae). Mammalian Species 48 (939): 91-108, DOI: 10.1093/mspecies/sew010, URL: http://dx.doi.org/10.1093/mspecies/sew010

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Additional details

Identifiers

LSID
urn:lsid:zoobank.org:pub:2846ACD3-C2D2-4E6F-B136-2A5B50BF04AA
LSID
urn:lsid:plazi.org:pub:0306FFE4FFEC7324BC79FC6FFFECDB4F
URL
http://publication.plazi.org/id/0306FFE4FFEC7324BC79FC6FFFECDB4F

References

  • ALLEN, J. A. 1974. On geographical variation in color among North American squirrels with a list of the species and varieties of the American Sciuridae occurring north of Mexico. Proceedings of the Boston Society of Natural History Volume 14:276-296.
  • ALVAREZ- CASTAnEDA, S. T., G. ARNAUD, and E. YENSEN. 1996. Spermophilus atricapillus. Mammalian Species 521:1-3.
  • ALVAREZ- CASTAnEDA, S. T., and P. CORTeS- CALVA. 2011. Genetic evaluation of the Baja California rock squirrel Otospermophilus atricapillus (Rodentia: Sciuridae). Zootaxa 3138:35-51.
  • BAKER, R. O. 1984. Commingling of Norway and roof rats with native rodents. Pp. 103-111 in Proceedings of the Eleventh Vertebrate Pest Conference (D. O. Clark, ed.). University of California, Davis.
  • BARBOUR, M. A., and R. W. CLARK. 2012. Ground squirrel tail-flag displays alter both predatory strike and ambush site selection behaviors of rattlesnakes. Proceedings of the Royal Society B 279:3827-3833.
  • BAYOUMI, A., and M. A. MEGUID. 2011. Wildlife and safety of earthen structures: a review. Journal of Failure Analysis and Prevention 11:295-319.
  • BEDNEKOFF, P. A., and D. T. BLUMSTEIN. 2009. Peripheral obstructions influence marmot vigilance: integrating observational and experimental results. Behavioral Ecology 20:1111-1117.
  • BEECHEY, F. W. 1941. An account of a visit to California 1926-'27. Reprinted from a narrative of a voyage to the Pacific and Beering's Strait performed in His Majesty's Ship Blossom under the command of Captain F. W. Beechey in 1825, '26, '27. '28. Grabhorn Press, San Francisco, California.
  • BIARDI, J. E., R. G. COSS, and D. G. SMITH. 1999. California ground squirrel (Spermophilus beecheyi) blood sera inhibits crotalid venom proteolytic activity. Toxicon 38:713-721.
  • BIARDI, J. E., D. C. CHIEN, and R. G. COSS. 2005. California ground squirrel (Spermophilus beecheyi) defenses against rattlesnake venom digestive and hemostatic toxins. Journal of Chemical Ecology 31:2501-2518.
  • BLACK, C. C. 1963. A review of the North American Tertiary Sciuridae. Bulletin of the Museum of Comparative Zoology 130:109-248.
  • BLOIS, J. L., R. S.FERANEC, and E.A.HADLY. 2008. Environmental influences on spatial and temporal patterns of body-size variation in California ground squirrels (Spermophilus beecheyi). Journal of Biogeography 35:602-613.
  • BLUMSTEIN, D. T., L. K.CHUNG, and J. E.SMITH. 2013. Early play may predict later dominance relationships in yellow-bellied marmots (Marmota flaviventris). Proceedings of the Royal Society of B 280:20130485.
  • BOELLSTORFF, D. E., and D. H. OWINGS. 1995. Home range, population structure, and spatial organization of California ground squirrels. Journal of Mammalogy 76:551-561.
  • BOELLSTORFF, D. E., D. H. OWINGS, M. C. T. PENEDO, and M. J. HERSEK. 1994. Reproductive behaviour and multiple paternity of California ground squirrels. Animal Behaviour 47:1057-1064.
  • BOYCE, D.A., J R. 1985. Prairie falcon prey in the Mojave Desert, California. Raptor Research 19:128-134.
  • BRYANT, W. E. 1889. Provisional description of supposed new mammals from California and Lower California. Proceedings of the California Academy of Sciences 2:25-27.
  • BURSTEN, S. N., K. C. BERRIDGE, and D. H. OWINGS. 2000. Do California ground squirrels (Spermophilus beecheyi) use ritualized syntactic cephalocaudal grooming as an agonistic signal? Journal of Comparative Psychology 114:281.
  • BURSTEN, S. N., R. B. KIMSEY, and D. H. OWINGS. 1997. Ranging of male Oropsylla montana fleas via male California ground squirrel (Spermophilus beecheyi) juveniles. Journal of Parasitology 83:804-809.
  • CARLTON, J. T., and J. HODDER. 2003. Maritime mammals: terrestrial mammals as consumers in marine intertidal communities. Marine Ecology Progress Series 256:271-286.
  • CARNIE, S. K. 1954. Food habits of nesting golden eagles in the coast ranges of California. Condor 56:3-12.
  • CLARK, J. P. 1994. Acrolein as a ground squirrel fumigant. Pp. 78-79 in Proceedings of the 16th Vertebrate Pest Conference (W. S. Halverson and A. C. Crabb, eds.). University of California, Davis.
  • CLUCAS, B., D. H. OWINGS, and M. P. ROWE. 2008b. Donning your enemy's cloak: ground squirrels exploit rattlesnake scent to reduce predation risk. Proceedings of the Royal Society B 275:847-852.
  • CLUCAS, B., M. P. ROWE, D. H. OWINGS, and P. C. ARROWOOD. 2008a. Snake scent application in ground squirrels, Spermophilus spp.: a novel form of antipredator behaviour? Animal Behaviour 75:299-307.
  • COLLINS, P. W., N. F. R. SNYDER, and S. D. EMSLIE. 2000. Faunal remains in California condor nest caves. Condor 102:222-227.
  • COPPETO, S. A., D. A. KELT, D. H. VAN VUREN, J. A. WILSON, and S. BIGELOW. 2006. Habitat associations of small mammals at two spatial scales in the northern Sierra Nevada. Journal of Mammalogy 87:402-413.
  • COSS, R. G. 1991. Context and animal behavior III: the relationship of early development and evolutionary persistence of ground squirrel antisnake behavior. Ecological Psychology 3:277-315.
  • COSS, R. G., and J. E. BIARDI. 1997. Individual variation in the antisnake behavior of California ground squirrels (Spermophilus beecheyi). Journal of Mammalogy 73:294-310.
  • COSS, R. G., K. L. GUSE, N. S. PORAN, and D. G. SMITH. 1993. Development of antisnake defenses in California ground squirrels (Spermophilus beecheyi): II. Microevolutionary effects of relaxed selection from rattlesnakes. Behaviour 124:137-164.
  • COSS, R. G., and D. H. OWINGS. 1978. Snake-directed behavior by snake naive and experienced California ground squirrels in a simulated burrow. Zeitschrift fur Tierpsychology 48:421-435.
  • CUVIER, G. 1831. Supplement a l'histoire naturelle generale et particuliere de Buffon, Tome I, Mammiferes. D. Pillot, Paris, France.
  • CYPHER, B. L., K. A. SPENCER, and J. H. SCHRIVNER. 1996. Use of food items by sex and age classes of coyotes. California Fish and Game 82:42-47.
  • DAAR/IPM. 2006. Integrated pest management plan for ground squirrels. City and County of San Francisco. DAAR/IPM Consulting Group, San Francisco, California. http://cchp.ucsf.edu/sites/cchp.ucsf.edu/files/ GroundSquirrels_FCCH_IPM.pdf. Accessed 14 November 2016.
  • DAVIDSON, A. D., J. K.DETLING, and J. H. BROWN. 2012. Ecological roles and conservation challenges of social, burrowing, herbivorous mammals in the world's grasslands. Frontiers in Ecology and the Environment 10:477-486.
  • DE SZALAY, F., L. C. CARROLL, J. A.B EAM, and V. H. RESH. 2003. Temporal overlap of nesting duck and aquatic invertebrate abundances in the Grasslands Ecological Area, California, U.S.A. Wetlands 23:739-749.
  • DIXON, J. S. 1906. Land birds of San Onofre, California. Condor 8:91-98.
  • DOBSON, F. S. 1979. An experimental study of dispersal in the California ground squirrel. Ecology 60:1103-1109.
  • DURANT, P., J. W. DOLE, and G. F. FISLER. 1988. Agonistic behavior of the California ground squirrel, Spermophilus beecheyi, at an artificial food source. Great Basin Naturalist 48:19-24.
  • EASTMAN, L. M., T. L. MORELLI, K. C. ROWE, C. J. CONROY, and C. MORITZ. 2012. Size increase in high elevation ground squirrels over the last century. Global Change Biology 18:1499-1508.
  • EDER, T., and G.ROSS. 2005. Mammals of California. Lone Pine Publishing, Auburn, Washington.
  • ELLIOT, D. G., et al. 1904. Descriptions of apparently new species and subspecies of mammals and a new generic name proposed. Field Columbian Museum, Publication 90. Zoological Series 3:263-270.
  • EMLEN, J. T., JR., and B. GLADING. 1938. California ground squirrel robs nest of valley quail. Condor 40:41-42.
  • ERXLEBEN, J. C. P. 1777. Systema Regni Animalis. Vol. 1. Impensis Weygandianis, Lipsiae, Germany.
  • ESTEP, J. A. 1989. Biology, movements, and habitat relationships of the Swainson's hawk in the Central Valley of California, 1986-87. California Department of Fish and Game, Nongame Bird and Mammal Section Report, Davis.
  • EVANS, F. C., and R. HOLDENRIED. 1943. A population study of the Beecheyi ground squirrel in central California. Journal of Mammalogy 24:231-260.
  • EVERMANN, B. W. 1882. American barn owl. Ornithologist and Oologist 7:97-98.
  • FEHMI, J. S., S. E. RUSSO, and J. W. BARTOLOME. 2005. The effects of livestock on California ground squirrels (Spermophilus beecheyii). Rangeland Ecology and Management 58:352-359.
  • FISHER, A. K. 1893. The hawks and owls of the United States in their relation to agriculture. U.S. Department of Agriculture, Division of Ornithology and Mammalogy, Washington Government Printing Office, Washington, D. C.
  • FITCH, H. S. 1947. Predation by owls in the Sierra foothills of California. Condor 49:137-151.
  • FITCH, H. S. 1948. Ecology of the California ground squirrel on grazing lands. American Midland Naturalist 39:513-596.
  • FITCH, H. S. 1949. Study of snake populations in central California. American Midland Naturalist 41:513-579.
  • FITCH, H. S., F. SWENSON, and D. F. TILLOTSON. 1946. Behavior and food habits of the red-tailed hawk. Condor 48:205-237.
  • FOWLER, F. H. 1931. Studies of food and growth of the prairie falcon. Condor 33:193-201.
  • FURMAN, D. P., and E. C. LOOMIS. 1984. The ticks of California (Acari: Ixodida). University of California Press, Berkeley.
  • GASHWILER, J. S. 1970. Plant and mammal changes on a clearcut in westcentral Oregon. Ecology 51:1018-1026.
  • GILSON, A., and T. P. SALMON. 1990. Ground squirrel burrow destruction: control implications. Pp. 97-98 in Proceedings of the 14th Vertebrate Pest Conference (L. R. Davis and R. E. Marsh, eds.). University of California, Davis.
  • GOLIGHTLY, R. T., JR., M. R. FAULHABER, K. L. SALLEE, and J. C. LEWIS. 1994. Food habits of introduced red fox in southern California. Pp. 15-20 in Proceedings of the Sixteenth Vertebrate Pest Conference (W. S. Halverson and A. C. Crabb, eds.). University of California, Davis.
  • GRINNELL, J. 1933. Review of the recent mammal fauna of California. University of California Publications in Zoology 40:71-234.
  • GRINNELL, J., and J. S. DIXON. 1918. Natural history of the ground squirrels of California. California State Printing Office, Sacramento.
  • GRINNELL, J., J. S. DIXON, and J. M. LINSDALE. 1937. Fur-bearing mammals of California. University of California Press, Berkeley.
  • GRINNELL, J., and T. I. STORER. 1924.Animal life in the Yosemite: an account of the mammals, birds, reptiles, and amphibians in a cross section of the Sierra Nevada. University of California Press, Berkeley.
  • HALL, E. R. 1926. Changes during growth in the skull of the rodent Otospermophilus grammurus beecheyi. University of California Publications in Zoology, University of California Press 21:355-404.
  • HALL, E. R. 1927. A commensal relation of the California quail and the California ground squirrel. Condor 29:271.
  • HALL, E. R. 1946. Mammals of Nevada. University of California Press, Berkeley.
  • HALL, E. R. 1981. The mammals of North America. 2nd ed. John Wiley & Sons, Inc., New York.
  • HANSON, M. T., and R. G. COSS. 1997. Age differences in the response of California ground squirrels (Spermophilus beecheyi) to avian and mammalian predators. Journal of Comparative Psychology 111:174-184.
  • HANSON, M. T., and R. G. COSS. 2001. Age differences in arousal and vigilance in California ground squirrels (Spermophilus beecheyi). Developmental Psychobiology 39:199-206.
  • HELGEN, K. M., F. R. COLE, L. E. HELGEN, and D. E. WILSON. 2009. Generic revision in the Holarctic ground squirrel genus Spermophilus. Journal of Mammalogy 90:270-305.
  • HENNESSY, D. F., and D. H. OWINGS. 1978. Snake species discrimination and the role of olfactory cues in the snake-directed behavior of the California ground squirrel. Behaviour 65:115-124.
  • HENNESSY, D. F., D. H. OWINGS, M. P. ROWE, R. G. COSS, and D. W. LEGER. 1981. The information afforded by a variable signal: constraints on snake-elicited tail flagging by California ground squirrels. Behaviour 78:188-226.
  • HERRON, M. D., T. A. CASTOE, and C. L. PARKINSON. 2004. Sciurid phylogeny and the paraphyly of Holarctic ground squirrels (Spermophilus). Molecular Phylogenetics and Evolution 31:1015-1030.
  • HOLDENRIED, R., F. C. EVANS, and D. S. LONGANECKER. 1951. Host-parasitedisease relationships in a mammalian community in the central coast range of California. Ecological Monographs 21:1-18.
  • HOLEKAMP, K. E., and S. NUNES. 1989. Seasonal variation in body weight, fat, and behavior of California ground squirrels (Spermophilus beecheyi). Canadian Journal of Zoology 67:1425-1433.
  • HOLEKAMP, K. E., S. NUNES, and F.TALAMANTES. 1988. Patterns of progesterone secretion in free-living California ground squirrels (Spermophilus beecheyi). Biology of Reproduction 39:1051-1059.
  • HOWARD, W. E., and J. R. BENTLEY. 1959. Competition between ground squirrels and cattle for range forage. Journal of Range Management 12:110-115.
  • HOWELL, A.H. 1931. Preliminary descriptions of four new North American ground squirrels. Journal of Mammalogy 12:160-162.
  • HOWELL, A. H. 1938. Revision of the North American ground squirrels with a classification of the North American Sciuridae. North American Fauna 56:1-256.
  • HUBBART, J. A., D. S. JACHOWSKI, and D. A. EADS. 2011. Seasonal and among-site variation in the occurrence and abundance of fleas on California ground squirrels (Otospermophilus beecheyi). Journal of Vector Ecology 36:117-123.
  • HUEY, L. M. 1931.Two new ground squirrels from Lower California, Mexico. Transactions of the San Diego Society of Natural History 7:17-20.
  • INGLES, L. G. 1945. Observations on the tree-climbing habits of California ground squirrels. Journal of Mammalogy 26:438.
  • JACHOWSKI, D. S., and J. A. HUBBART. 2012. Seasonal variation in pelage coloration of California ground squirrels (Otospermophilus beecheyi) in areas of differing land use. Journal of Biology and Life Sciences 3:13-19.
  • JACOBSEN, W. C. 1918. A history of ground squirrel control in California. The Monthly Bulletin, California State Commission of Horticulture 7:721-761.
  • JAEGAR, E. C. 1955. A source-book of biological names and terms. 3rd ed. Charles C. Thomas Publisher, Springfield, Illinois.
  • JEFFERSON, G. T. 1991. A catalogue of late quaternary vertebrates from California: Part 2: Mammals. Natural History Museum of Los Angeles County Technical Report 7.
  • JOHNS, D. W., and K. B. ARMITAGE. 1979. Behavioral ecology of alpine yellow-bellied marmots. Behavioral Ecology and Sociobiology 5:133-157.
  • KLAUBER, L. M. 1972. Rattlesnakes, their habits, life histories, and influence on mankind. 2nd ed. University of California Press, Berkeley.
  • KOENIG, W. D., and R. L. MUMME. 1987. Population ecology of the cooperatively breeding acorn woodpecker. Monographs in Population Biology, Princeton University Press, Princeton, New Jersey.
  • KOFORD, C. B. 1953. The California condor. National Audubon Society Research Report 4:1-154.
  • KRYGER, Z., L. GALLI- RESTA, G. H. JACOBS, and B. E. REESE. 1998. The topography of rod and cone photoreceptors in the retina of the ground squirrel. Visual Neuroscience 15:685-691.
  • KUENZI, A. J. 1997. Diet of coyotes at Lemoore Naval Air Station. Transactions of the Western Section of the Wildlife Society 33:9-11.
  • KURTEN, B., and E. ANDERSON. 1980. Pleistocene mammals of North America. Columbia University Press, New York.
  • LANG, J. D. 2004.Rodent-flea-plague relationships at the higher elevations of San Diego County, California. Journal of Vector Ecology 29:236-247.
  • LANG, J. D., and W. WILLIS. 1991. Ecology of sylvatic plague in the San Jacinto Mountains of southern California. Bulletin of the Society for Vector Ecology 16:183-199.
  • LAYMON, S. A. 1988. The ecology of the spotted owl in the central Sierra Nevada, California. Ph.D. dissertation, University of California, Berkeley.
  • LEGER, D. W., and D. H. OWINGS. 1978. Responses to alarm calls by California ground squirrels: effects of call structure and maternal status. Behavioral Ecology and Sociobiology 3:177-186.
  • LENIHAN, C. M. 2007. The ecological role of the California ground squirrel (Spermophilus beecheyi). Ph.D. dissertation, University of California, Davis.
  • LEOPOLD, S. A. 1977. The California quail. University of California Press, Berkeley.
  • LINSDALE, J. M. 1931. Facts concerning the use of thallium in California to poison rodents. Condor 33:96-106.
  • LINSDALE, J. M. 1946. The California ground squirrel: a record of observations made on the Hastings natural history reservation. University of California Press, Berkeley.
  • LITTRELL, E. E. 1990. Effects of field vertebrate pest control on nontarget wildlife (with emphasis on bird and rodent control). Pp. 59-61 in Proceedings of the Fourteenth Vertebrate Pest Conference (L. R. Davis and R. E. Marsh eds.). University of California, Davis.
  • LOREDO, I., D. VAN VUREN, and M. L. MORRISON. 1996. Habitat use and migration behavior of the California tiger salamander. Journal of Herpetology 30:282-285.
  • LOREDO- PRENDEVILLE, I., D.VAN VUREN, A. J. K UENZI, and M. M. MORRISON. 1994. California ground squirrels at Concord Naval Weapons Station: alternatives for control and the ecological consequences. Pp. 72-77 in Proceedings of the Sixteenth Vertebrate Pest Conference (W. S. Halverson and A. C. Crabb, eds.). University of California, Davis.
  • MARION, P. L., et al. 1987. Ground squirrel hepatitis virus and hepatocellular carcinoma. Pp. 337-348 in Hepadnaviruses (W. S. Robinson, K. Koike, and H. Will). Alan R. Liss, New York.
  • MARSH, R. E. 1994. Current (1994) ground squirrel control practices in California. Pp. 61-65 in Proceedings of the Sixteenth Vertebrate Pest Conference (W. S. Halverson and A. C. Crabb, eds.), University of California, Davis.
  • MARSH, R. E. 1998. Historical review of ground squirrel crop damage in California. International Biodeterioration and Biodegradation 42:93-99.
  • MARSH, R. E., R. H. SCHMIDT, and W. E. HOWARD. 1987. Secondary hazards to coyotes of ground squirrels poisoned with 1080 or strychnine. Wildlife Society Bulletin 15:380-385.
  • MCGRANN, M. C., D. H. VAN VUREN, and M. A. ORDEnANA. 2014. Influence of adjacent crop type on occurrence of California ground squirrels on levees in the Sacramento Valley, California. Wildlife Society Bulletin 38:111-115.
  • MEARNS, E. A. 1907. Mammals of the Mexican boundary of the United States: a descriptive catalogue of the species of mammals occurring in that region; with a general summary of the natural history, and a list of trees. Government Print Office 56: Smithsonian Institution, United States National Museum. http://wildsonora.com/sites/default/files/ reports/mammals-of-the-mexican-boundary-of-the-united-statesmearns-1907.pdf. Accessed 14 November 2016.
  • MERRIAM, C. H. 1893. Descriptions of eight new ground squirrels of the genera Spermophilus and Tamias from California, Texas, and Mexico. Biological Society of Washington Proceedings 8:129-138.
  • MERRIAM, C. H. 1910. The California ground squirrel. U. S. Bureau Biological Survey Circular 76:1-15.
  • MIAN, L. S., C. N. NWADIKE, J. C. HITCHCOCK, M. B. MADON, and C. M. MYERS. 1996. Plague activity in San Bernardino County during 1995. Proceedings of the Mosquito and Vector Control Association of California 64:80-84.
  • MILLER, A. H., and R. C. STEBBINS. 1964. The lives of desert animals in Joshua Tree National Monument. University of California Press, Berkeley.
  • MILLER, W. E. 1978. Pleistocene vertebrates of the Los Angeles basin and vicinity (exclusive of Rancho La Brea). Bulletin of the Los Angeles County Museum of Natural History 10:1-124.
  • MITROVICH, M. J., and R. A. COTRONEO. 2006. Use of plasticine replica snakes to elicit antipredator behavior in the California ground squirrel (Spermophilus beecheyi). Southwestern Naturalist 51:263-267.
  • MOORE, C. M., and P. W. COLLINS. 1995. Urocyon littoralis. Mammalian Species 489:1-7.
  • MURDOCH, J. D., K. RALLS, and B. L. CYPHER. 2004. Two observations of tree climbing by the San Joaquin kit fox. Southwestern Naturalist 49:522-523.
  • NADLER, C. F. 1966. Chromosomes of Spermophilus franklinii and taxonomy of the ground squirrel genus Spermophilus. Systematic Zoology 15:199-206.
  • NAVIN, T. R., et al. 1985. Human and sylvatic Trypanosoma cruzi infection in California. American Journal of Public Health 75:366-369.
  • NIETO, N. C., et al. 2007. Ectoparasite diversity and exposure to vectorborne disease agents in wild rodents in central coastal California. Journal of Medical Entomology 44:328-335.
  • NOWICKI, S., and K. B. ARMITAGE. 1979. Behavior of juvenile yellow-bellied marmots: play and social integration. Zeitschrift fur Tierpsychologie 51:85-105.
  • NUSSBAUM, R. A., and C. MASER. 1975. Food habits of the bobcat, Lynx rufus, in the Coast and Cascade ranges of western Oregon in relation to present management policies. Northwest Science 49:261-266.
  • OAKS, E. C., P. J. YOUNG, G. L. KIRKLAND, JR., and D. F. SCHMIT. 1987. Spermophilus variegatus. Mammalian Species 272:1-8.
  • O'CONNELL, R. A. 1994. Trapping ground squirrels as a control method. Pp. 66-67 in Proceedings of the Sixteenth Vertebrate Pest Conference (W. S. Halverson and A. C. Crabb, eds.). University of California, Davis.
  • OLSON, A. C., JR. 1950. Ground squirrels and horned larks as predators upon grunion eggs. California Fish and Game 36:323-327.
  • ORDEnANA, M. A., D. H. VAN VUREN, and J. P. DRAPER. 2012. Habitat associations of California ground squirrels and Botta's pocket gophers on levees in California. Journal of Wildlife Management 76:1712-1717.
  • ORLOFF, S., F. HALL, and L. SPIEGEL. 1986. Distribution and habitat requirements of the San Joaquin kit fox in the northern extreme of their range. Transactions Western Section of the Wildlife Society 22:60-70.
  • OWINGS, D. H., M. B. BORCHERT, and R. VIRGINIA. 1977. The behaviour of California ground squirrels. Animal Behaviour 25:221-230.
  • OWINGS, D. H., and R. G. COSS. 1977. Snake mobbing by California ground squirrels: adaptive variation and ontogeny. Behaviour 62:50-69.
  • OWINGS, D. H., and R. G. COSS. 2007. Hunting California ground squirrels: constraints and opportunities for northern Pacific rattlesnakes. Pp. 155-167 in Biology of the rattlesnakes (W. K. Hayes, K. R. Beaman, M. D. Cardwell, and S. P. Bush, eds.). Loma Linda University Press, Loma Linda, California.
  • OWINGS, D. H., D. F. HENNESSY, D. W. LEGER, and A. B. GLADNEY. 1986. Different functions of "alarm" calling for different time scales: a preliminary report on ground squirrels. Behaviour 99:101-116.
  • OWINGS, D. H., and D. W. LEGER. 1980. Chatter vocalizations of California ground squirrels: predator- and social-role specificity. Zeitschrift fur Tierpsychology 54:163-184.
  • PAJAK, A. F., III, E. SCOTT, and C. J. BELL. 1996. A review of the biostratigraphy of Pliocene and Pleistocene sediments in the Elsinore Fault Zone, Riverside County, California. Pp. 28-49 in The uses of vertebrate fossils in biostratigraphic correlation (C. J. Bell and S. S. Sumida, eds.). University of California, Berkeley.
  • PATTEE, O. H., P. H. BLOOM, J. M. SCOTT, and M. R. SMITH. 1990. Lead hazards within the range of the California condor. Condor 92:931-937.
  • PATTEN, M. A. 1997. Reestablishment of a rodent community in restored desert scrub. Restoration Ecology 5:156-161.
  • PHUONG, M. A., M. C. W. LIM, D. R. WAIT, K. C. ROWE, and C. MORITZ. 2014. Delimiting species in the genus Otospermophilus (Rodentia: Sciuridae), using genetics, ecology, and morphology. Biological Journal of the Linnean Society 113:1136-1151.
  • PORAN, N. S., and R. G. COSS. 1990. Development of antisnake defenses in California ground squirrels (Spermophilus beecheyi): I. Behavioral and immunological relationships. Behaviour 112:222-245.
  • PORAN, N. S., R. G. COSS, and E. BENJAMIN. 1987. Resistance of California ground squirrels (Spermophilus beecheyi) to the venom of the northern Pacific rattlesnake (Crotalus viridis oreganus): a study of adaptive variation. Toxicon 25:767-777.
  • PURCELL, K. L., and J. VERNER. 1999. Nest predators of open and cavity nesting birds in oak woodlands. Wilson Bulletin 111:251-256.
  • PUTMAN, B. J., and R. W. CLARK. 2015. The fear of unseen predators: ground squirrel tail flagging in the absence of snakes signals vigilance. Behavioral Ecology 26:185-193.
  • RABIN, L. A., R. G. COSS, and D. H. OWINGS. 2006. The effects of wind turbines on antipredator behavior in California ground squirrels (Spermophilus beecheyi). Biological Conservation 131:410-420.
  • RANCK, R. S., J. M. CULLEN, K. S. WAGGIE, and P. L. MARION. 2008. Haderian gland neoplasms in captive, wild-caught Beecheyi ground squirrels (Spermophilus beecheyi). Veterinary Pathology 45:388-392.
  • RICHARDSON, J. 1829. Fauna Boreali-Americana of the zoology of the northern parts of British America. London, United Kingdom.
  • ROBERTS, S. L., D. A. KELT, J. W. VAN WAGTENDONK, A. K. MILES, and M. D. MEYER. 2015. Effects of fire on small mammal communities in frequent-fire forests in California. Journal of Mammalogy 96:107-119.
  • RODRIGUEZ- ROBLES, J. 2002. Feeding ecology of North American gopher snakes (Pituophis catenifer, Colubridae). Biological Journal of the Linnean Society 77:165-183.
  • ROEST, A. I. 1993. Ground squirrels feeding on eelgrass. California Fish and Game 79:85-86.
  • ROWE, M. P., and D. H. OWINGS. 1978. The meaning of the sound of rattling by rattlesnakes to California ground squirrels. Behaviour 66:252-267.
  • ROWE, M. P., and D. H. OWINGS. 1990. Probing, assessment, and management during interactions between ground squirrels and rattlesnakes. Part 1: risks related to rattlesnake size and body temperature. Ethology 86:237-249.
  • ROWE, M. P., and D. H. OWINGS. 1996. Probing, assessment and management during interactions between ground squirrels (Rodentia: Sciuridae) and rattlesnakes (Squamata: Viperidae). 2: cues afforded by rattlesnake rattling. Ethology 102:856-874.
  • RUNDUS, A. S., D. H. OWINGS, S. S. JOSHI, E. CHINN, and N. GIANNINI. 2007. Ground squirrels use an infrared signal to deter rattlesnake predation. Proceedings of the National Academy of Sciences 104:14372-14376.
  • RUTLEDGE, L. C., M. A. MOUSSA, B. L. ZELLER, and M. A. LAWSON. 1979. Field studies of reservoirs and vectors of sylvatic plague at Fort Hunter Liggett, California. Journal of Medical Entomology 15:452-458.
  • RYAN, R. M. 1968. Mammals of Deep Canyon, Colorado Desert, California. The Desert Museum, Palm Springs, California.
  • RYCKMAN, R. E., and C. C. LINDT. 1953. Aberrant Citellus beecheyi specimens from Orange County, California. Journal of Mammalogy 34:266.
  • SALMON, T. P. 1981. Artificial establishment of a ground squirrel colony. Journal of Wildlife Management 45:1016-1018.
  • SANDBERG, S., and B. H. BANTA. 1973. Instances of southern California ground squirrels (Spermophilus beecheyi nudipes) eating iguanid lizards. Herpeton 7:7-8.
  • SAVAGE, D. E. 1949. Pleistocene vertebrates of the San Francisco Bay region. Ph.D. dissertation, University of California, Berkeley.
  • SAVAGE, D. E., and D. E. RUSSELL. 1983. Mammalian paleofaunas of the world. Benjamin/Cummings, Menlo Park, California.
  • SAYLOR, L. W. 1937. Red-tailed hawk choked by squirrel skull. Condor 39:175-176.
  • SCHITOSKEY, F., JR. 1975. Primary and secondary hazards of 3 rodenticides on kit fox. Journal of Wildlife Management 39:416-418.
  • SCHITOSKEY, F., JR., and S. R.WOODMANSEE. 1978. Energy requirements and diet of the California ground squirrel. Journal of Wildlife Management 42:373-382.
  • SCHRAMM, R., and G. BULLARD. 2004. Williams and Associates, Inc. Fresh Carrot Advisory Board of California. Comment 1015 submitted to the United States Environmental Protection Agency Docket OPP-2004-0033.
  • SHAPIRO, M. R., et al. 2010. Rickettsia 364D: a newly recognized cause of eschar-associated illness in California. Clinical Infectious Diseases 50:541-548.
  • SIMPSON, G. L., and T. K. LAMUNYON. 1980. Multiple litters in the California ground squirrel, Spermophilus beecheyi fisheri, in Tulare County. Pp. 101-104 in Proceedings of the 9th Vertebrate Pest Conference. Digital Commons at the University of Nebraska-Lincoln, Lincoln.
  • SORK, V. L. 2016. Gene flow and natural selection shape spatial patterns of genes in tree populations: implications for evolutionary processes and applications. Evolutionary Applications 9:291-310
  • SPANO, R. K. 1994. Ground squirrel management in the Angeles National Forest. Pp. 68-71 in Proceedings of the Sixteenth Vertebrate Pest Conference (W. S. Halverson and A. C. Crabb, eds.). University of California, Davis.
  • STANTON, F. W. 1944. Douglas ground squirrel as a predator on nests of upland game birds in Oregon. The Journal of Wildlife Management 8:153-161.
  • STOCK, C. 1972. Rancho La Brea.A record of Pleistocene life in California, no. 20. 6th ed. Los Angeles County Museum of Natural History Science Series, Los Angeles, California.
  • STONER, E. A. 1949. Butcher bird feeds on ground squirrel. Wilson Bulletin 64:187.
  • STORER, T. I. 1930. Summer and autumn breeding in the California ground squirrel. Journal of Mammalogy 11:235-237.
  • STROUD, D. C. 1982. Dispersal and some implications for control of the California ground squirrel. Pp. 210-213 in Proceedings of the Tenth Vertebrate Pest Conference (R. E. Marsh, ed.). University of California, Davis.
  • SWAISGOOD, R. R., M. P. ROWE, and D. H. OWINGS. 1999. Assessment of rattlesnake dangerousness by California ground squirrels: exploitation of cues from rattling sounds. Animal Behaviour 57:1301-1310.
  • SWAISGOOD, R. R., M. P. ROWE, and D. H. OWINGS. 2003. Antipredator responses of California ground squirrels to rattlesnakes and rattling sounds: the roles of sex, reproductive parity, and offspring age in assessment and decision-making rules. Behavioral Ecology and Sociobiology 55:22-31.
  • TIETJE, W. D., S. L. NIVES, J. A.HONIG, and W. H. WEITKAMP. 1991. Effect of acorn planting depth on depredation, emergence, and survival of valley and blue oak. United States Department of Agriculture Forest Service General Technical Report Pacific Southwest 126:14-20.
  • THOMSON, G. M. 1922. The naturalisation of animals and plants in New Zealand. Cambridge University Press, London, United Kingdom.
  • THORINGTON, R. W., JR., and R. S. HOFFMANN. 2005. Family Sciuridae. Pp. 754-818 in Mammal species of the world: a taxonomic and geographic reference (D. E. Wilson and D. M. Reeder, eds.). Johns Hopkins University Press, Baltimore, Maryland.
  • THORINGTON, R. W., JR., J. L. KOPROWSKI, M. A. STEELE, and J. F. WHATTON. 2012. Squirrels of the world. John Hopkins University Press, Baltimore, Maryland.
  • TIMM, R. M., R. O. BAKER, J. R. BENNETT, and C. C. COOLAHAN. 2004. Coyote attacks: an increasing suburban problem. 69th North American Wildlife and Natural Resources Conference 69:67-88.
  • TOMICH, P. Q. 1962. The annual cycle of the California ground squirrel Citellus beecheyi. University of California Publication in Zoology 65:213-282.
  • TOWELL, D. E., and R. G. ANTHONY. 1988. Annual diet of bobcats in Oregon's Cascade Range. Northwest Science 62:99-103.
  • TOWNZEN, K. R., M. A. THOMPSON, and C. R. SMITH. 1996. Investigations and management of epizoic plague at Ice House Reservoir, Eldorado National Forest, California, 1994 and 1995. Pp. 68-74 in Proceedings of the 14th Vertebrate Pest Conference (L. R. Davis and R. E. Marsh, eds.), University of California, Davis.
  • TRENHAM, P. C., and H. B. SHAFFER. 2005. Amphibian upland habitat use and its consequences for population viability. Ecological Applications 15:1158-1168.
  • VANDER HAEGAN, W. M., G. R. ORTH, and L. M. AKER. 2005. Ecology of the western gray squirrel in south-central Washington. Washington Department of Fish and Wildlife, Olympia.
  • VAN VUREN, D. H., A. J. KUENZI, I. LOREDO, A. L. LEIDER, and M. L. MORRISON. 1997. Translocation as a nonlethal alternative for managing California ground squirrels. The Journal of Wildlife Management 61:351-359.
  • VAN VUREN, D. H., and M. A. ORDEnANA. 2012. California levee vegetation research program: burrow dimensions of ground squirrels, with special reference to the California ground squirrel. Proceedings of the 2012 California Levee Vegetation Research Program Symposium. University of California, Davis.
  • VON BLOEKER, J. C. 1967. Land mammals of the southern California islands. Pp. 245-263 in Proceedings of the Symposium on the biology of the California islands (R. W. Philbrick, ed.). Santa Barbara Botanic Gardens, Santa Barbara, California.
  • VON BLOEKER, J. C., and R. L. RUDD. 1937. Two unusual screech owls. Condor 39:176.
  • WAGGIE, K. S., and P. L. MARION. 1997. Demodex sp. in California ground squirrels. Journal of Wildlife Diseases 33:368-370.
  • WARREN- ALLEN, R. A. 2004. Road crossings: a survey of wildlife presence and mortality along roads in agricultural Oregon. M.S. thesis, Willamette University, Salem, Oregon.
  • WEINTRAUB, J. D. 1986. Coyote diets, five years later, at Cuyamaca Rancho State Park. Bulletin of the Southern California Academy of Sciences 85:152-157.
  • WELLMAN, C., and W. B. WHERRY. 1910. Some new internal parasites of the California ground squirrel (Otospermophilus beecheyi). Parasitology 3:417-422.
  • WHERRY, W. B. 1908. Plague among the ground squirrels of California. Journal of Infectious Disease 5:485-506.
  • WHISSON, D. A. 1999. Modified bait stations for California ground squirrel control in endangered kangaroo rat habitat. Wildlife Society Bulletin 27:172-177.
  • WHITE, P. J., K. RALLS, and C. A. VANDERBILT WHITE. 1995. Overlap in habitat and food use between coyotes and San Joaquin kit foxes. Southwestern Naturalist 40:342-349.
  • WINCHELL, C. S. 1994. Natural history and protection of burrowing owls. Pp. 83-86 in Proceedings of the Sixteenth Vertebrate Pest Conference (W. S. Halverson and A. C. Crabb, eds.). University of California, Davis.
  • WOBESER, G. A. 1994. Disease management through manipulation of the host population Pp. 153-167 in Investigation and management of disease in wild animals (G. A. Wobeser, ed.). Springer, New York.
  • YEH, P. J., M. E. HAUBER, and T. D. PRICE. 2007. Alternative nesting behaviors following colonization of a novel environment by a passerine bird. Oikos 116:1473-1480.
  • ZIELINSKI, W. J., N. P.DUNCAN, E. C.FARMER, R. L.TRUEX, A. P.C LEVENGER, and R. H. BARRETT. 1999. Diet of fishers (Martes pennanti) at the southernmost extent of their range. Journal of Mammalogy 80:961-971.