ATYPENA FORMOSANA (OI, 1977)

(FIGS 7H, 67I, 68I, 73)

Callitrichia formosana Oi, 1977: 23, figs 1–5 (Dmf).

Oedothorax formosanus Brignoli, 1983: 349.

Callitrichia formosana Song, 1987: 144, fig. 104 (Tmf from Oedothorax).

Atypena formosana Tazoe, 1992: 212, figs 1–10 (mf; NB: generic placement following Jocqué, 1983).

Oedothorax formosanus Okuma et al., 1993: 13, fig. 11A–B (f).

Atypena formosana Barrion & Litsinger, 1994: 319, figs 1673–1676 (mf).

Callitrichia formosana Song et al., 1999: 160, fig. 88K–L (mf).

Callitrichia formosana Ono et al., 2009: 267, figs 95–99 (mf).

Examined material: Japan: Okinawa, Iromotejima Island, Urauchi, 2♂ 3♀ 02.iv.1995, leg. A. Tanikawa, det. A, Tanasevitch (SMF 56485).

Diagnosis:

Males: This species can be distinguished from A. cirrifrons by the proportionally shorter distal suprategular apophysis and ventral radical process.

Females: No study of Atypena comparative female morphology exists to date; such diagnosis is, therefore, beyond the scope of the current study.

Description:

Male (SMF): Total length: 1.76. Prosoma: 0.78 long, 0.62 wide, PME region largely elevated, interocular region with translucent setae laterally curved (Fig. 7H). Eyes: AME-AME: 0.03, AME width: 0.06, AME- ALE: 0.05, ALE width: 0.08, ALE-PLE: 0.01, PLE width: 0.08, PLE-PME: 0.14, PME width: 0.07, PME- PME: 0.17. Clypeus: upper-half hirsute, one sub- AME seta. Sternum: 0.46 long, 0.46 wide. Chelicerae: mastidia absent; stridulatory striae imbricated, rows compressed proximally (Fig. 67I). Legs: tibia chaetotaxy 2-2-1-1, dorsal proximal macroseta on tibia I, II, III and IV 0.65, 0.82, 1.45 and 1.87 times diameter of tibia, respectively; Tm I: 0.78. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPA short, pointed and sclerotized at tip; PC base not visible from dorsal view, distal setae close to distal clasp, terminal part slightly enlarged, distal clasp without striae, clasp extended apically (Fig. 73A); T without papillae; PT small, without papillae; TS long and slender, without papillae (Fig. 73D); MSA absent; DSA wide, truncated at tip, ventral corner more extended than dorsal corner (Fig. 73A); EM broad and flat, without papillae, exceeding ARP; ARP flat, unsclerotized; LER retrolateral to E; VRP long, slender; R without papillae; TP long, slender; E short and stout, slightly retrolaterally curved (Fig. 73B). Opisthosoma: dorsal pattern see Fig. 68I; PMS with mAP, three AC; PLS with triad, 3+ AC.

Female (SMF): See also Oi (1977); epigyne without prominent external modification, copulatory ducts slightly curved, trojactory simple, entrances to spermathecae mesal to exits of fertilixation ducts. (Fig. 73). Opisthosoma: dorsal pattern same as male; PMS with CY, mAP, 2–4 AC; PLS with triad, 2 CY, 3+ AC (Fig. 73).

Distribution: Bangladesh to Japan.

Remarks: As in A. cirrifrons, a scaly sac-like structure is observed at the paracymbial base (Figs 73A, C, 74A, C). According to the results of our phylogenetic analysis, and the similarities in male genital organs and prosomal modifications, this species is congeneric with A. cirrifrons.

ADDITIONAL MORPHOLOGICAL DATA