(Figs. 24, 196)

Description. Occiput black, frons dark brown to reddish brown posteriorly, orange anteriorly, rest of head orange. Prementum and maxillary palp yellow. Mostly covered with microtomentum, frons with shiny spots lateral to ocelli and a small spot between ocellar bristles, face with a small ventromedial shiny patch. Ocellar bristles just anterior to median ocellus. Subvibrissal and anterior genal bristles about 0.3X length of vibrissa. Gena height about 0.4x height of eye.

Thorax black, postpronotum, posterior corners of scutum, and a band from base of wing to mid coxa dark reddish brown. Mostly covered with microtomentum, proepisternum shiny, anepisternum with a large ventromedial shiny spot covering about 0.3X length and 0.5X height. Halter whitish, brown below knob.

Legs dark brown, joints, trochanters, and tarsi orange, fore coxa reddish brown. Mid tibia with row of anterodorsal, 1 anteroventral, 1 posteroventral, 5 subapical bristles. Hind tibia with a thin anteroventral, 3 ventroapical bristles.

Wing light brown, veins brown, crossveins r-m and dm-cu white. Vein R 4+5 with faint indication of 2 paler spots distal to crossvein r-m.

Abdominal tergites heavily sclerotized, black, covered with microtomentum. Sternites heavily sclerotized, black, wider than long, covered with microtomentum in male, weakly sclerotized, longer than wide in female. Pleural setae long and pliant.

Male postabdomen: Sternite 5 with curved lateral margins, widest near apex, anterior apodeme broad, about 1.6X length of external portion. Surstylus scooped. Pregonite distinct. Postgonite with lobes well separated, anterior lobe short and rounded, posterior lobe longer and pointed. Basiphallus with short broad epiphallus and preepiphallus, preepiphallus transparent. Distiphallus with curved spinose dorsal tube flanked by two spike-like sclerites.

Female postabdomen: Tergites 6–7 wider than long, weakly sclerotized medially, covered with microtomentum except anterior margin. Sternites 6–7 moderately sclerotized, covered with microtomentum except anterior margin. Tergites with 3 posterior strips, sternites with 2 posterior strips. Tergite 8 approximately square, with anterior and posterior margins shallowly emarginated, covered with microtomentum except anteromedial patch. Epiproct with long anterior arms, 1 pair of setae, distal portion with microtomentum. Cerci thin. Sclerites of sternite 8 broad, pinched near base, distal half covered with microtomentum. Hypoproct subrectangular, with a curved unsclerotized band medially, covered with microtomentum except anterior margin. Spermathecae barrelshaped, slightly longer than wide, with long basal invagination, sclerotized duct about 1.5X as long as spermathecae.

Type material. Holotype ♂: ECUADOR: Napo: Lago Papallacta, nr., 3400 m, forest above lake, pans/ dung, 4–8 Nov 1999, S.A. Marshall, debu00135682 (QCAZ). Paratypes: same data as holotype (2 ♂, 4 ♀, DEBU); Quito–Baeza pass, 1 km E, 3950 m, forest edge, dung traps, 4–8 Nov 1999, S.A. Marshall (1 ♀, DEBU); Quito– Baeza road, Papallacta, 3200 m, elfin forest, above thermal spgs., pan traps, 16–21 Feb 1983, L. Masner (1 ♀, DEBU).

Comments. This species is similar to A. variegatus sp. nov., but can be separated based on the pattern of microtomentum on the face and anepisternum. The species name is derived from the Greek ano- (above) and the Latin variegatus, referring to the varicoloured thorax and the altitudinal distribution relative to A. variegatus.