Geron senilis Fabricius
Authors/Creators
Description
Geron senilis (Fabricius)
Bombylius senilis Fabricius, 1794: 411. Turton 1801: 683. Fabricius 1805: 135. Wiedemann 1821: 171.
Geron senilis (Fabricius). Wiedemann 1828: 357. Macquart 1846: 119. Osten Sacken 1858: 43. Osten Sacken 1877: 264. Osten Sacken 1878: 94. Coquillett 1894: 111. Johnson 1894: 275. Aldrich 1905: 245. Johnson 1908: 72. Kertész 1909: 99. Johnson 1910: 747. Johnson 1913: 58. Walton 1915: 173. Cockerell 1917: 14. Sturgis 1918: 28 (as “ semlis ”). Johnson 1919: 429. Cole & Lovett 1921: 251. Painter 1932: 144, 165, 166. Priddy 1939: 47. Painter & Painter 1962: 18. Hull 1973: 204, 206. Painter, Painter & Hall 1978: 17. Evenhuis 1983: 488. Scarbrough & Davidson 1985: 1241, 1242. Evenhuis & Greathead 1999: 48. Godinez Alvarez 2003: 24.
Geron albaria Painter, 1932: 145, 159. Byers et al. 1962: 143. Tabet 1974: 20. Tabet & Hall 1984: 32.
Geron albarius Painter. Painter & Painter 1965: 422 (emendation). Hull 1973: 206. Miller 1978: 42. Frommer 1981: 35. Evenhuis 1983: 462. Tabet & Hall 1987: 67. Poole & Gentili 1996: 70.
Bombylius insularis Bigot, 1857a:332 (1857b: 792).
Geron insularis (Bigot). Osten Sacken 1878: 94. Aldrich1905: 245. Kertész 1909: 99. Painter & Painter 1962: 47. Painter, Painter & Hall 1978: 17. Evenhuis 1983: 476. Scarbrough & Davidson 1985: 1242.
Geron antologos Tabet, 1979: 185. Evenhuis 1983: 463. Nomen nudum.
Geron antologos Tabet, 2001: 49. Syn. nov.
Description. Male. Head. Front silvery tomentose, pile usually wanting; antenna black; scape and pedicel white pilose; first flagellomere linear, evenly tapering from base to acuminate apex; antennal ratio: 9:3:30; face cinereous pollinose, bare medially, silvery tomentum opposite antennal base; oral margin white; proboscis extends beyond oral margin for a distance approximately two times head height; gena white pilose and tomentose; occiput white pilose, silvery tomentum on lower half next to eye margin, wanting on upper half; ocellar tubercle with dark yellow hair.
Thorax. Mesonotum velvety black, side and anterior margin lightly cinereous pollinose; median longitudinal stripe brownish gray, short, broad, not divided, reaches transverse suture; mesonotal pile white to pale yellow anteriorly, dark yellow to black posteriorly, tomentum golden yellow, sparse; pleura white pilose and tomentose; legs black, mid tibia ferruginous, legs with white hair and scales; halter stem brownish; knob white; scutellum with pale yellow hair and golden yellow tomentum.
Wing. Hyaline; venation normal; crossvein rm at middle of cell dm; vein mcu sinuous; basicosta with pale yellow hair.
Abdomen. Dorsum white to pale yellow pilose, golden yellow tomentose; side and venter white pilose and tomentose; lateral margins of tergites and posterior margins of sternites narrowly pale.
Genitalia (Fig. 53) in ventral view with gonocoxa flat with two illdefined lateral, posteriorly converging ridges, median ridge (when present) not prominent; gonocoxa in lateral view rectangular, lower margin without projections; gonostylus long, narrow, four times longer than high; lateral ramus narrow, shorter than gonostylus; dorsolateral ramus short, about half length of lateral ramus.
Female. Eyes separated at vertex by approximately one and onehalf times width of ocellar tubercle; front with pale yellow hair, tomentum yellow medially, silvery next to eye margin and immediately above antennae; occiput with yellow pile and golden yellow tomentum on upper half; ocellar tubercle with dark yellowish hair. Thorax. Mesonotum cinereous, median longitudinal stripe narrow, not divided, reaching beyond transverse suture; notum white pilose anteriorly, yellow pilose posteriorly, tomentum golden yellow, abundant; legs black, mid and hind tibiae brownish; fore tarsi with palynophilic setae; scutellum with yellow hair and golden yellow tomentum. Abdomen. Dorsum yellow pilose, golden yellow tomentose. Genitalia with posterior projection on tergite VIII black; vaginal furca (Fig. 22) usually heavily sclerotized, upper half narrowly Vshaped, lower half stout, lateral arms thick, tapering from narrowed base to outwardly curved apex, inner margin at apex swollen, giving a somewhat bilobed appearance, without internal rami.
Types. Lectotype male of Bombylius senilis (“ America meridionalis insularis ”) in the Lund and Sehestedt Collection, ZMUC designated by Scarbrough & Davidson (1985: 1244); not seen by authors. Lectotype male of G. insularis (Cuba) in MNHN studied and designated by Painter & Painter (1962: 47); seen by junior author. Holotype male of G. albaria (Brownwood, Texas, 18.vii.1921, R.H. Painter) and allotype female (Brownwood, Texas, 23.vii.1921, Painter) both in USNM (No. 44869). Holotype male of G. antologos (12 mi. H. Gooding, Gooding Co., Idaho, 11.viii.1977, on Grindelia squamosa, A.B. Tabet) in CAS (No. 14296). Types of G. albaria studied by senior author.
Material Examined. United States. Arizona. Pima County: 2, Molino basin, Santa Catalina Mtns., milepost 4 Hitchcock Hwy, 29.iv.1961 (R.H. & E.M. Painter) (USNM). California. Kern County: 1, Freeman, 1.viii.1956 (T.R. Haig) (UCR). Inyo County: 1, 12 mi. E. Keeler, 7.vii.1982 (J.C. & E.M. Painter) (UCR); 13, Lone Pine, Diaz Lake, 14.vi.1972 (J.D. Pinto); 4, 1 mi. N. Lone Pine, 12.vi.1964 (M.E. Irwin) (UCR). Modoc County: 18, 5 mi. E. Newell, 17.vii.1980 (J.C. & E.M. Hall) (UCR). Riverside County: 1, Whitewater Canyon, 11.ix.1935 (P.H. Timberlake) (UCR); 12, Santa Rosa Mtn., 2.vi.1987 (J.C. Hall) (UCR); 1, Smith Water Canyon, 30.viii.1986 (K.W. Cooper) (UCR). San Bernardino County: 6, Morongo Valley, 5.x.1934 (A.L. Melander) (USNM). Idaho. Bonneville County: 5, 11 mi. W. Idaho Falls, 7.viii.1958 (R.H. & E.M. Painter). Illinois. 1, [southern] (Robertson) (USNM). Maryland. Montgomery County: 1, Plummer’s Island, 20.vii.1913 (W.L. McAtee) (USNM). Montana. Madison County: 1, Gardiner, 20.viii.1918 (A.L. Melander) (USNM). Missoula County: 1, Missoula, 16.viii.1904 (USNM). Nevada. Lyon County: 2, 25 mi. NE Fernley, 29.viii.1956 (T.R. Haig) (UCR). Washoe County: 1, 2 mi. N. Nixon, 16.ix.1972 (J.D. Pinto) (UCR); 1, Patrick, 16.vi.1964 (A.G. Gillogly) (UCR). Pershing County: 1, 20 mi. S. Lovelock, 30.viii.1957 (R.W. Lauderdale) (UCR). New Jersey. Gloucester County: 2, Malaga, viii.1940 (G.M. Greene) (USNM). New Mexico. Hidalgo County: 3, 7 mi. SE Rodeo, 21.viii.1958 (R.E. Rice, G.B. Pitman, D.D. Linsdale) (UCB). Nevada. Lyon County: 6, 25 mi. NE Fernley, 29.viii.1956 (T.R. Haig) (UCR). Utah. Iron County: 2, Anderson’s Wash, 29.vi.1919 (USNM); Cane County: 3, Bryce Canyon N.P. 27.viii.1957 (R.H. & E.M. Painter) (USNM). Virginia. Fairfax County: Mt. Vernon, 4.vii.1917, on flowers of Xolisma ligustrina (W.L. McAtee) (USNM). Washington. Grant County: 4, Stratford, 4.ix.1920 (R.C. Shannon) (USNM). Garfield County: 1, Wawai, 30.viii.1908 (N.M. Mann) (USNM). Douglas County: 1, Wonatchoo, 8.vi.1919 (A.L. Melander) (USNM). Mexico. Durango: 4, Los Nieves, 8700 ft., 5.ix.1962 (R.H. & E.M. Painter) (USNM). Oaxaca: 3, 4 mi. E. Salina Cruz, 7.ix.1963 (R.H. & E.M. Painter) (USNM). San Luis Potosí: 2, 40 mi. N. San Luis Potosí, 8.viii.1963 (R.H. & E.M. Painter (USNM). Veracruz: Cordoba, reared from larval case of Tineola uterella Walsingham, adult issued 2.iv.04 (F. Knab) (USNM). Jamaica. 3, South West Point, 23.viii.1984, dry scrub (N.E. Woodley) (USNM).
Variation. Nearly all of the diagnostic characteristics are subject to variation, however, the majority of the specimens studied have yellow tomentum on the front of the head and along the dorsum of the body. In the male genitalia the venter of the gonocoxa may have two shallow depressions not separated by a definite ridge. There appears to be considerable variation in the development of the apical lobes and in the degree of sclerotization. The apical lobes vary from slightly swollen on the lower inner margin, to bilobed, to greatly swollen on the lower inner margin. In general, specimens within local populations are usually quite uniform as to the shape of the vaginal furca, but there appears to be too much interpopulation variation and overlap to be able to consider treating the species listed in the synonymy above as subspecies.
Discussion. This species is much like G. albidipennis. The male genitalia show only slight differences, the two depressions and the strong median dividing ridge are more prominent in G. albidipennis. The vaginal furca in the female of G. senilis has the lower half not as stoutly developed as in G. albidipennis and the spine on the lower surface is less developed. The vaginal furca also exhibits some resemblance to that of G. argutus. In the latter species the apex of the lower half of the apodeme is bent outward, footlike, and there are no swellings or lobes.
Evenhuis & Greathead (1999) synonymized G. antologos (then a nomen nudum) under G. senilis based on unpublished notes in this manuscript regarding similar configuration of the male genitalia. The subsequent valid publication of G. antologos by Tabet (2001) requires it to be formally synonymized here.
Biology. One single female specimen in the USNM collection has labels stating that it was reared from the larval case of Tineola uterella Walsingham from Cordoba, Mexico. There is no associated pupal case pinned with this specimen. Adults have been taken at flowers of Xolisma ligustrina, Stipulicida setacea, Grindelia squamosa, Chaenactis douglasii and Salsola sp. Tabet (1974) hypothesized that because of the pattern of collections of this species (as G. albaria) in Deep Canyon, California, it might exhibit trivoltinism.
Distribution. United States (Arizona, California, Colorado, Florida, Idaho, Illinois, Kansas, Maryland, Montana, Nevada, New Jersey, New Mexico, Texas, Utah, Virginia, Washington, Wyoming); Mexico (Durango, Nuevo León, Oaxaca, Puebla, San Luis Potosí, Veracruz); Bahamas; Cuba; Jamaica; U. S. Virgin Is; Puerto Rico.
Notes
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Linked records
Additional details
Identifiers
Biodiversity
- Scientific name authorship
- Fabricius
- Kingdom
- Animalia
- Phylum
- Arthropoda
- Order
- Diptera
- Family
- Bombyliidae
- Genus
- Geron
- Species
- senilis
- Taxon rank
- species
References
- Fabricius, J. C. (1794) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tome IV. C. G. Proft, Hafniae [= Copenhagen], [6] + 472 + [5] pp.
- Turton, W. (1801) A general system of nature, through the three grand kingdoms, of animals, vegetables, and minerals: systematically divided into their several classes, orders, genera, species, and varieties, with their habitations, manners, economy, structure and peculiarities. Translated from Gmelins's last edition of the celebrated Systema Naturae, by Sir Charles Linne: amended and enlarged by the improvements and discoveries of later naturalists and societies, with appropriate copper-plates. Vol. III. " 1800 " Lackington, London, 784 pp.
- Fabricius, J. C. (1805) Systema antliatorum secundum ordines, genera, species adiecta synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunsvigae [= Brunswick], xiv + [15] - 372 + [1] + 30 pp.
- Wiedemann, C. R. W. (1821) Diptera exotica. [Ed. 2]. Pars I. Tabulis aeneis duabus. Kiliae [= Kiel], xix + 244 pp.
- Wiedemann, C. R. W. (1828) Aussereuropaische zweiflugelige Insekten. Als Fortsetzung des Meigenschen Werkes. Erster Theil. Schulz, Hamm, xxxii + 608 pp.
- Macquart, P. J. M. (1846) Dipteres exotiques nouveaux ou peu connus. Supplement. N. E. Roret, Paris. Pp. 5 - 238. [This is the reprint version issued in advance of the journal article, 1846, Mem. Soc. R. Sci. Agric. Arts, Lille 1844: 133 - 364 and presented by Macquart at the 22 July 1846 meeting of the Societe Entomologique de France. The date on the wrapper of the journal is 1846, the date on the title page is 1845. The last 4 pages of this paper in the journal (containing the index) are numbered 363, 364, 363, 364. The next article in the journal begins on page 365. The pagination of the reprint is continuous, thus a direct comparison of the journal and reprint pagination causes the reprint to appear to contain 2 additional pages. The journal version recorded in the 7 November 1846 issue of Bibliogr. Fr.]
- Osten Sacken, C. R. (1858) Catalogue of the described Diptera of North America. Smithsonian Miscellaneous Collections, 3 (1), 1 - 87.
- Osten Sacken, C. R. (1877) Western Diptera: descriptions of new genera and species of Diptera from the region west of the Mississippi and especially from California. Bulletin of the United States Geological Survey of the Territories, 3 (2), 1 - 354.
- Osten Sacken, C. R. (1878) Catalogue of the described Diptera of North America. [Second edition.]. Smithsonian Miscellaneous Collections, 270, 1 - 276.
- Coquillett, D. W. (1894) Notes and descriptions of North American Bombyliidae. Transactions of the American Entomological Society, 21, 89 - 112.
- Johnson, C. W. (1894) List of the Diptera of Jamaica, with descriptions of new species. Proceedings of the Academy of Natural Sciences of Philadelphia, 46, 271 - 281.
- Aldrich, J. R. (1905) A catalogue of North American Diptera (or two-winged flies). Smithsonian Miscellaneous Collections, 46 (1444), 1 - 680.
- Johnson, C. W. (1908) The Diptera of the Bahamas, with notes and description of one new species. Psyche, 15, 69 - 80.
- Kertesz, K. (1909) Catalogus dipterorum hucusque descriptorum. Volumen V. Bombyliidae, Therevidae, Omphralidae. G. Engelmann, Budapest, 199 pp.
- Johnson, C. W. (1910) Order Diptera, pp. 703 - 814. In: Smith, J. B., Insects of New Jersey. Annual Report of the New Jersey State Museum, 1909, 15 - 888.
- Johnson, C. W. (1913) Insects of Florida. I. Diptera. Bulletin of the American Museum of Natural History, 32, 37 - 90.
- Walton, W. R. (1915) Report on some parasitic and predaceous Diptera from northeastern New Mexico. Proceedings of the United States National Museum, 48 (2070), 171 - 186.
- Cockerell, T. D. A. (1917) The fauna of Boulder County, Colorado, III. Class Insecta, Order Diptera. University of Colorado Studies, 12, 5 - 20.
- Sturgis, W. (1918) Claremont-Laguna Diptera from the collections of the Department of Zoology of Pomona College. Journal of Entomology and Zoology, 10, 27 - 31.
- Johnson, C. W. (1919) A revised list of the Diptera of Jamaica. Bulletin of the American Museum of Natural History, 41 (8), 421 - 449.
- Cole, F. R. & Lovett, A. L (1921) An annotated list of the Diptera (flies) of Oregon. Proceedings of the California Academy of Sciences, (4) 11, 197 - 344.
- Painter, R. H. (1932) A monographic study of the genus Geron as it occurs in the United states (Diptera: Bombyliidae). Transactions of the American Entomological Society, 58, 139 - 167.
- Priddy, R. B. (1939) List of Bombyliidae collected in southern California and Yuma County, Arizona. Journal of Entomology and Zoology, 31, 45 - 53.
- Painter, R. H. & E. M. Painter. (1962) Notes on and redescriptions of types of North American Bombyliidae (Diptera) in European Museums. Journal of the Kansas Entomological Society, 35, 1 - 164.
- Hull, F. M. (1973) Bee flies of the world. The genera of the family Bombyliidae. Bulletin of the United States National Museum, 286, 1 - 687.
- Painter, R. H., Painter, E. M. & Hall, J. C. (1978) Family Bombyliidae. A catalog of the Diptera of the Americas south of the United States, 38, 1 - 92.
- Evenhuis, N. L. (1983) An indexed bibliography of Bombyliidae (Insecta, Diptera). Theses Zoologicae, 4, 1 - 493.
- Scarbrough, A. G. & D. A. Davidson (1985) Review of the Caribbean Geron Meigen (Diptera: Bombyliidae). Journal of the New York Entomological Society, 93 (4), 1240 - 1260.
- Evenhuis, N. L. & Greathead, D. J. (1999) Catalog of the bee flies of the world (Diptera: Bombyliidae). Backhuys Publishers, Leiden, xlviii + 756 pp.
- Godinez Alvarez, A. (2003) Bombyliidae (Diptera) de las Reservas Especiales de la Biosfera de Ria Celestun y de Ria Lagartos, Yucatan, Mexico. Unpublished Master's thesis, Universidad Autonoma de Mexico, Mexico, D. F, 64 pp. + appendices and plates.
- Byers, G. W., Blank, F., Hanson, W. J., Beneway, D. F. & Fredrickson, R. W. (1962) Catalogue of the types in the Snow Entomological Museum. Part III (Diptera). University of Kansas Science Bulletin, 43, 131 - 181.
- Tabet, A. B. (1974) A phenology study of the Bombyliidae of Deep Canyon. Unpublished Master's Degree thesis, University of California, Riverside, viii + 94 pp.
- Tabet, A. B. & J. C. Hall. (1984) The Bombyliidae of Deep Canyon. Part I. A phenology study of the Bombyliidae of Deep Canyon. Al-Fateh University Publications, Tripoli, Libya, 63 pp.
- Painter, R. H. & Painter, E. M. (1965) Family Bombyliidae. In: Stone, A., Sabrosky, C. W, Wirth, W. W., Foote, R. H. & Coulson, J. R .. (eds.), A catalog of the Diptera of America north of Mexico. United States Department of Agriculture, Agriculture Handbook, 276, pp. 406 - 46.
- Miller, E. L. (1978) A checklist of the Diptera in the South Dakota State University insect collection. I. Suborders Nematocera and Brachycera. Proceedings of the South Dakota Academy of Science, 57, 32 - 45.
- Frommer, S. I. (1981) A hierarchic listing of the arthropods known to occur in the environs of the P. L. Boyd Deep Canyon Desert Research Center. Philip Boyd Deep Canyon Desert Research Center, University of California, Natural Land and Water Reserves System, xv + 124 pp.
- Tabet, A. B. & J. C. Hall. [1987] The Bombyliidae of Deep Canyon. Part II. " 1984 ". Al-Fateh University Publications, Tripoli, Libya. 176 pp. [The date on the cover is " 1984 " (the same as the publication date for Part I of this work). It was not received by Jack Hall until the summer of 1987 and his resulting correspondence with Tabet confirmed that this part was not issued until March 1987.]
- Poole, R. W. & Gentili, P. (editors) (1996) Nomina Insecta Nearctica. A checklist of the insects of North America. Volume 3: Diptera, Lepidoptera, Siphonaptera. Entomological Information Services, Rockville, Maryland, 1143 pp.
- Bigot, J. M. F. (1857 a) Dipteros. In: Sagra, R. de la, Historia fisica politica y natural de la Isla de Cuba. Segunda parte. Historia natural. Tomo VII. Crustaceos, aragnides e insectos. A. Bertrand, Paris, pp. 328 - 49.
- Tabet, A. B. (1979) The bee flies of Idaho. Unpublished Ph. D. dissertation, University of Idaho, Moscow, xxii + 595 pp.
- Tabet, A. B. [2001] Three new species of Geron from Idaho (Diptera: Bombyliidae). Journal of Basic and Applied Sciences (Tripoli), 6 [1998], 48 - 52. [The date on the cover is " 1998 ", but correspondence with Tabet confirmed that this part was not issued until June 2001.]