(Figs. 228–230, 363, Map 42)

Diagnosis.— Glyphidocera speculae is most similar to G. vestita but differs from the latter by having an abdomen with more rows of sex scales between terga 2–3 in male, a tegumen that is narrower basally, wider costal furcae of valva, a more angular apicoventral margin of cucullus, and an apical cornutus without a bulbous base, and the absence of within vesica.

Description.— Head: Vertex and frontoclypeus pale orange intermixed with few paleyellow scales; outer margin of labial palpus with segments I–II brown intermixed with pale­orange scales to apical margin, segment III brown with few pale­orange scales on apical half; inner surface pale orange intermixed with few brown scales; scape of antenna pale orange intermixed with few brown scales, flagellum with alternating bands of brown and pale­orange scales; male 4 th flagellomere dorsally protuberant and curved, forming a deep concavity opposite slightly excavated flagellomeres 5–6; concavity nearly closed by a few setiform scales on apex of protuberance of 4 th flagellomere; proboscis pale orange.

Thorax: Tegula brown basally, pale orange distally; mesonotum pale orange. Legs pale yellow intermixed with pale­orange and few brown scales, pale yellow to apical margin of all segments and tarsomeres. Forewing (Fig. 363): Length 6.9 mm (n = 1), pale orange intermixed with pale­yellow scales, and brown scales along costa and from near distal end of cell to margin. Undersurface brown. Hindwing: Pale brown.

Abdomen (Fig. 228): Male with 5 transverse, irregular rows of sex scales on intersegmental area between terga 2–3.

Male Genitalia (Figs. 229–230): Uncus elongate, wide at base, broadly constricted at 1 / 3, slightly rounded distally, apex narrowly rounded and setose; gnathos sparsely setose, protuberant, wide basally, gradually narrowing apically, upturned apically; tegumen narrow, slightly widened basally; dorsal strut of tegumen with arms convergent basally and dorsally, forming an elliptical­shaped sinus between points of constriction, dorsally atrophied; valva with costa slightly sloped from a shallow depression beyond a slanted, basal, digitate process to dorsal furca; apex widely bifurcate, furcae slightly divergent, dorsal furca large, falcate, ventral furca extending slightly beyond outer margin of cucullus; sacculus twisted apically, constricting valva near midlength, widening distoventrally, forming an elongate and slightly angular cucullus; cucullus membranous, setose; vesica denticulate, bearing a moderately large, hornlike, apical cornutus; cornutus with one side slightly shortened, exposing a hollow, basal cavity.

Female Genitalia: Unknown.

Holotype, ɗ, “Est[ación] Sirena, P[arque] N[acional] Corcovado, Prov[incia] Punta[renas], COSTA RICA, 0–100 m, Nov. 1993, G. Fonseca, L­S­ 270500, 508300, # 2490 ”, “ INBio: COSTA RICA: CRI 001, 625857 ” [bar code label], “ INBio, ɗ Genitalia Slide by D. Adamski, No. 1086 ” [yellow label], “ PARATYPE, Glyphidocera guaroa, Adamski ” [blue label].

Distribution (Map 42): Glyphidocera speculae is known from a single collecting site on the Osa Peninsula in south­eastern Costa Rica.

Etymology: The species epithet speculae is derived from the Latin “specula,” meaning watchtower.