(Figs. 117–119, 326, Map 22)

Diagnosis.— Glyphidocera lupae is most similar to G. ciatae but differs from the latter by having a spatulate gnathos, a shorter digitate process of the valva, a narrower valval cucullus, and a larger apical cornutus of the aedeagal vesica.

Description.— Head: Vertex and frontoclypeus pale brownish yellow; outer surface of labial palpus pale yellowish brown intermixed with few brown scales, outer surface pale yellowish brown; scape and flagellum of antenna pale yellowish brown; male 4 th flagellomere distally dilated, protuberant and curved, forming a deep concavity opposite deeply excavated flagellomeres 5–6; concavity nearly closed by setiform scales on apex of protuberance of 4 th flagellomere; proboscis pale brownish yellow.

Thorax: Tegula and mesonotum pale brownish yellow. Legs pale brownish yellow or pale brownish yellow intermixed with few brown scales. Forewing (Fig. 326): Length 6.1 –6.9 mm (n = 4), pale brownish yellow or pale brownish yellow intermixed with few brown scales; cell with 2 spots, 1 near middle, 1 near distal end [specimens with spots present or absent]; marginal spots present or absent. Undersurface pale brownish yellow intermixed with few brown scales. Hindwing: Pale brownish yellow.

Abdomen (Fig. 117): Male with 4 transverse, irregular rows of equal­sized sex scales on intersegmental membrane between terga 2–3 and terga 3–4.

Male Genitalia (Figs. 118–119): Uncus elongate, wide at base, parallelsided apically, apex slighted rounded and setose; gnathos sparsely setose, protuberant, wide basally, gradually narrowed to a spatulate apex; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing midway between posteroventral margin of tegumen, gradually widening to base of uncus; costa of valva nearly straight, apical furca falcate; base of costa bearing a slanted, digitate process; sacculus twisted apically, constricting valva slightly before midlength, widening distoventrally, forming an elongate and broadly rounded cucullus; cucullus membranous and setose; vesica of aedeagus denticulate, with several small spinelike cornuti, and bearing an elongate, conical, apical cornutus; cornutus with one side shortened near midlength, exposing a hollow, basal cavity.

Female Genitalia: Unknown.

Holotype, ɗ, “Monumento Nacional Guayabo, Prov[incia] Carta[go], COSTA RICA, 1100 m, 28 Set.– 21 Nov. 1994, G. Fonseca, L­N­ 217400, 570000, # 3285 ”, “ INBio: COSTA RICA: CRI 002, 024744” [bar code label], “ INBio, Genitalia Slide by D. Adamski, Sex ɗ, No. 523 ” [yellow label].

Paratypes (3 ɗ): 1 ɗ, Same label data as holotype except, “ Nov. 1994, # 3287 ”, “ CRI 002, 092549 ”, “Slide No. 519 ”; “Parque Nacional Braulio Carrillo, Estac. Carrillo, 700 m, San José, COSTA RICA, Julio 1990, I Curso Microlepidoptera”, “ CRI 000, 171159 ”, “Slide No. 843 ”, “ CRI 000, 171119 ”, “Slide No. 845 ”, (2 in INBio, 1 in USNM).

Distribution (Map 22): Glyphidocera lupae is known from two collecting sites in north­central Costa Rica along the Cordillera Central.

Etymology: The species epithet lupae is derived from the Latin “lupa,” meaning a shewolf.