(Figs. 2 a, 18–20, 293, Map 4)

Diagnosis.— Glyphidocera operae is most similar to G. orae but differs by having a larger gnathos, a dorsal strut of the tegumen that is atrophied dorsally, a curved digitate process of the valva, a narrower distal part of the valva, and a longer apical cornutus of the aedeagus.

Description.— Head: Vertex and frontoclypeus brown; outer surface of labial palpus with segments I–II brown intermixed with few pale grayish­yellow scales near bases and apices, segment III brown; apical area of segment II and entire length of segment III with raised scales (Fig. 2 a); inner surface with segment I and basal half of segment II grayish yellow, distal half of segment II and segment III brown; scape of antenna dark brown, flagellomeres brown; 4 th flagellomere dorsally protuberant and curved, forming a narrow concavity opposite slightly flattened flagellomeres 5–6, concavity nearly closed by a few setiform scales on apex of protuberance of 4 th flagellomere; proboscis pale grayish orange intermixed with few brown scales.

Thorax: Tegula and mesonotum dark brown. Legs dark brown intermixed with pale grayish­orange scales on apices of all segments and tarsomeres. Forewing (Fig. 293): Length 7.0 mm (n = 1), dark brown intermixed with few grayish­brown scales. Undersurface brown. Hindwing: Brown.

Abdomen (Fig. 18): Male with 7 transverse, irregular rows of minute sex scales on intersegmental area between terga 2–3 and 3 transverse, irregular rows of sex scales on intersegmental membrane between terga 3–4.

Male Genitalia (Figs. 19–20): Uncus elongate, wide at base, broadly constricted near 1 / 3, slightly rounded medially, apex narrowly rounded and setose; gnathos protuberant, widened basally, abruptly narrowed, forming a knoblike and setose apical process; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing medially along posteroventral margin; costa slightly rounded broadly from a shallow depression beyond digitate basal process to an attenuate apical furca; base of costa with a stalked, slightly downcurved digitate process; sacculus twisted apically, constricting valva near 1 / 3 length, forming an elongate cucullus; cucullus membranous and setose, with outer margin convergent to valval apex; vesica of aedeagus densely denticulate, bearing an elongate, conical cornutus; cornutus with one side shortened near midlength, exposing a hollow, basal cavity.

Female Genitalia: Unknown.

Holotype, ɗ, “Est[ación] Pitilla 700 m, 9 km S S[an]ta Cecilia, P[arque] N[acional] Guanacaste, Prov[incia] Guan[acaste], COSTA RICA, C. Moraga, Jun. 1991, L­N­ 330200, 380200 ”, “ INBio: COSTA RICA: CRI 000, 699973 ” [bar code label], “ INBio, ɗ Genitalia Slide by D. Adamski, No. 753 ” [yellow label].

Paratype, 1 ɗ, Same label data as above except, “P. Rios & C. Morgana, 10 Set– 22 Oct 1990 ”, “ CRI 000, 182166 ”, “Slide No. 752.”

Distribution (Map 4): Glyphidocera operae is known from one collecting site in northwestern Costa Rica along the Cordillera de Guanacaste.

Etymology: The species epithet operae is derived from the Latin “opera,” meaning task.