Coeloides abdominalis (Zetterstedt), 1840 (Figs. 1–4)

Bracon abdominalis Zetterstedt, 1840, Insecta Lapponica, 1: 398.

Coeloides abdominalis (Zetterstedt): Thomson, 1892, Opusc. Ent., 17: 1845; Yang, 1996, Parasitic wasps on bark beetles in China, 24.

Material studied. 1Ψ CHINA, Heilongjiang, Yichun, 16. vi. 1956, Shen Nong, No. 948363 (CAFB); 43ΨΨ 73ɗɗ, CHINA, Shaanxi, Gaixian city, 5. iv. 1984, Yang Zhong­qi and Wang Bin­hai, ex. Ips acuminatus Gyllenhal (CAFB).

Diagnosis: This species is similar to Coeloides ungularis Thomson, but can be separated from the latter by the key mentioned above. It has antennae with 33–40 segments; first flagellomere slightly protruding apically, and second flagellomere distinct flared apico­ventrally, slightly shorter than the first one; third flagellomere weakly protruding basally and apically; notauli of mesoscutum wide and shallow, absent posteriorly; veins 2­SR: SR1 of fore wing = 1: 2.5; first metasomal tergite with wide, smooth and shallow grooves postero­laterally; second metasomal tergite with narrow and deep longitudinal grooves laterally; and ovipositor sheath relatively long, 1.4 times as long as abdomen.

Biology: The known hosts are the larvae of Ips acuminatus Gyllenhal and Ips typographus Linnaeus on Korean pine, Tomicus pilifer Spessivtseff on Pinus koraiensis Siebold & Zuccarini in China, which belong to family Scolytidae (Coleoptera) (Yang, 1996). The other known hosts are the larvae of the following: Melanophila cyanea F. of the family Buprestidae (Coleoptera) (Filippenkova, 1971; Tobias, 1986; Tobias, 1976); Blastophagus minor Hartig (Filippenkova, 1971; Hedqvist, 1998; Mills, 1983; Tobias, 1986; Tobias, 1976), Blastophagus piniperda L. (Capek et al., 1982; Filippenkova, 1971; Haeselbarth, 1967; Hedqvist, 1998; Mills, 1983; Tobias, 1976; Tobias, 1986), Blastophagus piniperda Linnaeus on Pinus sylvestris L. (Herard & Mercadier, 1996), Hylesinus fraxini Panzer (Hedqvist, 1998), Hylurgops palliates Gyllenhal (Hedqvist, 1998), Ips cembrae Heer (Haeselbarth, 1967; Hedqvist, 1998; Mills, 1983), Ips sexdentatus Boerner (Arefin, 1984; Filippenkova, 1971; Hedqvist, 1998; Mills, 1983; Tobias, 1986; Tobias, 1976), Ips subelongatus Motschlsky (Tobias, 1986; Tobias, 1976), Ips typographus L. (Haeselbarth, 1967; Hedqvist, 1998; Mills, 1983), Ips typographus japonicus De Geer (Lawson et al., 1996), Phloeotribus scarabaeoides Bernad (Hedqvist, 1998), Pissodes notatus (F.) (Alauzet, 1987; Alauzet, 1982; Haeselbarth, 1967; Tobias, 1976; Tobias, 1986), Pissodes notatus on Pinus sylvestris L. (Kenis & Mills, 1994), Pissodes validirostris Sahiberg (Jakimavicius, 1976), Scolytus multistriatus (Marsham) (Haeselbarth, 1967; Hedqvist, 1998), Scolytus ratzeburgi Janson (Hedqvist, 1998), Scolytus rugulosus (Muller) (Hedqvist, 1998) of the family Scolytidae (Coleoptera) and Acanthocinus aedilis (Linnaeus) of family Corydalidae (Megaloptera).

Distribution: China (Heilongjiang and Shaanxi province), Norway, Finland, Germany, Russia, Sweden, Hungary, England, Czechoslovakia, Poland, France, Japan, Turkey and Denmark (Yang, 1996; Shenefelt, 1978)

Note: Yang (1996) once reported this species as a subspecies, Coeloides abdominalis orientalis Haeselbarth, parasiting Ips typographus L. and Ips cembrae Heer in Shaanxi province of China, but the subspecific name is not adopted in this study.