Cladophorus haiaensis sp. nov.

(Figs. 9, 19, 23, 39, 59–60)

Type material. Holotype, male. Papua New Guinea, Crater Mt., Haia, Supa, 06°40´S 145°03´E, 1070–1450 m (A00124, LMBC).

Diagnosis. C. haiaensis has a smaller body compared with other Cladophorus and it is the only uniformly black coloured species. The species represents the deepest split in the genus (Fig. 4) and the different shape of the phallus supports its position (Figs. 59–60). Although having slightly different genitalia, C. haiaensis is robustly placed in Cladophorus (BS 88%) and the morphological divergence is limited to genitalia, which are under strong sexual selection.

Description. Male. Body small, 10.0 mm long, uniformly dark brown to black (Fig. 39). Head small, with very small eyes, frontal interocular distance 2.37 times maximum diameter; antennae long, reaching beyond half of elytral length, flabellate, antennomere 3 with lamella attached to apical part of antennomere (Fig. 9), but its apical part missing in holotype, antennomeres 7–9 with very long lamellae (Fig. 19). Pronotum transverse, much wider basally, 1.57 times wider than long at midline, lateral margins slightly elevated, frontal margin slightly rounded, frontal angles obtuse but conspicuous, lateral margins straight, posterior angles sharply prominent, postero-lateral carinae weak in middle of disc and antero-lateral carinae weak towards frontal margin, median anterior carina welldeveloped (Fig. 23). Elytra flat, only slightly widened posteriorly, transverse costae weak, irregular, partly covered by pubescence, elytral areoles variable in shape (Fig. 39). Phallus very slender, widest basally, narrowed gradually to apex (Figs. 59–60).

Measurements. BL 10.0 mm, PL 1.12 mm, PW 1.67 mm, WH 1.06 mm, Edist 0.83 mm, Ediam 0.35 mm, LE 8.5 mm, antennomere 3: measurements unavailable.

Distribution. Eastern New Guinea, Chimbu province (Fig. 3).

Etymology. The epithet refers to the local name ‘Haia’, the type locality of the species.