Campostichomma manicatum Karsch, 1892

Figures 2–4, 14

Campostichomma manicatum KArsch, 1892: 296, pl. 12, FIg. 19 (FEmAlE hOlOtypE FrOm SrI LAnkA, dEpOsItEd In MusEum Für NAturkundE, BErlIn, nOt ExAmInEd). SImOn 1898A: 7; GrIswOld 1993: 7, FIgs 31–36, 75, 77; WSC 2017

Note. WE hAVE nOt ExAmInEd thE hOlOtypE, but hAVE ExAmInEd thE PArIs spEcImEns thAt wErE studIEd by LEhtInEn (1967) And dEscrIbEd by SImOn (1989A). BEcAusE LEhtInEn AlsO ExAmInEd thE hOlOtypE, wE AccEpt thE IdEntIty OF thE PArIs spEcImEns As bEIng cOnspEcIFIc wIth C. manicatum.

Diagnosis. Males of Campostichomma manicatum can be distinguished from those of other Ububidae by characters of the male palp including the palpal tibia being less than ½ the length of cymbium (Fig. 3A), the cymbium with deep groove retrolaterally (Fig. 4A, C) and the embolus elongate, thread-like, extending 360 ˚ around tegulum (Figs 3D, 4A). The females of C. manicatum can be distinguished from those of other Campostichomma species by the epigynum without teeth (Figs 3F, 4D), with a broad and slightly sclerotized epigynal plate, and a small median lobe plate, and by the vulva with one loop of the ascending duct, which is broad anteriorly (Figs 3G, 4E).

DesCription. Male (MNHN—CASENT9067600): Total length 7.30. Markings as in Fig. 2A, B, E. Carapace 3.60 long, 2.90 wide, 1.40 high; clypeus 0.14 high. Eye diameters: AME 0.12, ALE 0.18, PME 0.12, PLE 0.20. Chelicerae 1.30 long; sternum 1.60 long, 1.35 wide; labium 0.70 long; palpal coxae 1.15 long. Spination as in genus description. Leg measurements (Femur + Patella + Tibia + Metatarsus + Tarsus = [Total]): I: 2.40 + 1.20 + 2.30 + 2.30 + 1.50 = 9.70; II: 2.40 + 1.10 + 1.90 + 2.10 + 1.20 = 8.70; III: 2.30 + 0.95 + 1.60 + 2.20 + 1.20 = 8.25; IV: 2.80 + 1.00 + 2.00 + 3.30 + 1.45 = 10.55. Leg formula 4123. Palp: palpal tibia shorter than cymbial length, RTA elongated, RTA length longer than tibia, with pointed apex (Fig. 3A–D); cymbium tapering gradually to base, dorsal scopulate patch sparse; tegulum suboval (Fig. 4A); MA origin central, cup-shaped MA, oval, (Fig. 4A–B); base of conductor narrow and conductor fan-shaped (Fig. 4A–C); embolus elongated, unmodified, spiralling around outer margin of tegulum and gradually tapering to pointed apex (Fig. 4A–C).

Female (MNHN): Total length 8.30. Markings as in male and as in Fig. 2C, D, F. Carapace 3.40 long, 2.60 wide, 1.80 high; clypeus 0.08 high. Eye diameters: AME 0.12, ALE 0.18, PME 0.12, PLE 0.20. Chelicerae 1.50 long; sternum 1.60 long, 1.30 wide; labium 0.60 long, palpal coxae 1.30 long. Spination as in genus description. Leg measurements (Femur + Patella + Tibia + Metatarsus + Tarsus = [Total]): I: 2.20 + 1.10 + 1.80 + 1.80 + 1.00 = 7.90; II: 2.10 + 1.20 + 1.70 + 1.50 + 1.00 = 7.50; III: absent; IV: absent. Epigynum: median lobe projected posteriorly and truncated at apex; lateral lobes without teeth (Fig. 4D). Vulva with elongated and sinuous copulatory ducts; spermathecae small, with projections and small fertilization ducts (Fig. 4E).

Variation. Male (N= 3): total length 6.20–7.40; carapace length 3.30–3.70; femur I 2.20–2.50. Female (N= 2): total length 6.30–6.70; carapace length 3.50–3.55; femur I 2.10–2.20.

Material examined: SRI LANKA. Central Province, 17.5 km (11 miles) SSE Nuwara-Eliya, Horton Plains National Park, 1 male, CASENT9067600, 19–20.III.1962, 2042 m (6700 ft) a.s.l. (UMLS); Central Province, Kandy, Mathurata, 5 females, 2 males, CASENT9047632 (MNHN 16242).

Natural history. The holotype was collected in the late 19th century near Kandy and Nuwara Eliya, two localities located in the highlands of southern Sri Lanka in Central Province, at above 800 m elevation. Today this area is largely cultivated for tea plantations but there are remnants of montane forests, especially near Nuwara Eliya and at Horton Plains National Park, at elevations from 1800 to more than 2200 m. Specimens were collected in these areas in the mid 20th century. Collection data for the specimens collected at Horton Plains refer to “bush and meadow” with “scattered trees of Rhododendron remaining.” The specimens were collected at a stream near the edge of a remaining primary forest. (Brinck et al. 1971: 36).

Distribution. Known from the highlands of Central Province in Sri Lanka (Fig. 14).