Epeorus (Caucasiron) bicolliculatus Hrivniak, sp. nov.

Figures 1–48

Epeorus alpicola (Eaton, 1871): Türkmen & Kazancı (2015: 44) [partim] Epeorus sylvicola (Pictet, 1865): Türkmen & Kazancı (2015: 44, 45) [partim] Epeorus (Caucasiron) sp.: Martynov et al. (2016: 170)

Type material. Locality numbers (in square brackets) correspond to numbers on Fig. 51. Collectors of type material are listed in section Material and methods.

Holotype: female mature larva, [1.] GEORGIA: Autonomous Republic of Adjara (hereafter Adjara), vicinity of Chakhati village, Kintrishi Riv. (41°45'43"N 41°58'34"E), 325 m a.s.l., Martynov A.V. leg. 18– 20.4.2016 (preserved in 96% EtOH).

Paratypes: 1 female, 1 male imago, 1 male subimago (all reared from larvae and mounted on slides), 1 male, 6 female larvae (2 mounted on slide), same place and date as holotype [1.]; same place and date as holotype [1.]; 2 female larvae, GEORGIA, Adjara, vicinity of Chakhati village, Kintrishi Riv. (41°45'42"N 41°58'42"E; close to holotype locality), 325 m a.s.l., 6.6.2016; 12 female larvae (1 mounted on slide, 3 with only mouthparts mounted on slide), 1 male larva (mounted on slide), [2.] GEORGIA: Adjara, vicinity of Chakhati village, right tributary of Kintrishi Riv. (41°44'14"N 41°59'00"E), 380 m a.s.l., 19.4.2016; 1 male larva, [4.] GEORGIA: Imereti Region (hereafter Imereti), vicinity of Tskhomareti village, Jruchula Riv. (42°23'12.8"N 43°21'33.3"E), 505 m a.s.l., 23.6.2013; 4 male, 5 female larvae (1 mounted on slide), [5.] GEORGIA: Imereti, right tributary of Sairmegele Riv. (41°52'57.7"N 42°45'22.6"E), 1276 m a.s.l., 25.6.2013; 2 female larvae (1 mounted on slide), [8.] GEORGIA: Imereti, Sairmegele Riv. and its right tributary (41°52'45.2"N 42°45'32.2"E), 1285 m a.s.l., 25.6.2013; 1 female larva, [9.] GEORGIA: Imereti, app. 2 km S. Health Resort Sairme, Sairmegele Riv. (41°52'56"N 42°45'09"E), 1262 m a.s.l., 25.6.2013; 1 male, 2 female larvae (1 mounted on slide), GEORGIA: Adjara, vicinity of Chakhati village, right tributary of Kintrishi Riv. (41°45'23"N 41°58'41"E; close to holotype locality), 385 m a.s.l., 7.6.2013; 1 female larva, [12.] GEORGIA: Guria Region, Ozurgeti (41°54'29.64"N 42°03'52.44"E), 169 m a.s.l., 22.8.2009; 1 female larva, [13.] GEORGIA: Adjara, Zeda Erge village, near bridge on road Batumi –Akhaltsikhe, Jochostskali Riv. (right tributary of Tchorokhi Riv.) (41°34'02.04"N 41°41'16.20"E), 40 m a.s.l., 21.8.2009; 2 female larvae, [14.] GEORGIA: Adjara, Adzarisagmarti village, left tributary of Machakhlistskali Riv. (41°30'37.20"N 41°49'18.06"E), 207 m a.s.l., 20.8.2009; 1 female larva, [15.] TURKEY: Trabzon Province, vicinity of Hamşiköy village, right tributary of Maçka Riv. (40°40'45.36"N 39°26'55.74"E), 1460 m a.s.l., 29.6.2011; 1 female larva, [16.] GEORGIA: Adjara, Khala village, Kemerkobis Riv. (right tributary of Chekvistskali Riv.) (41°42'08.70"N 41°48'51.30"E), 100 m a.s.l., 18.8.2009; 5 female larvae (2 mounted on slides), 1 male larva (mounted on slide) [19.] TURKEY: Giresun Province, Kümbet Plateau, small tributary of Aksu Stream (40°32'19''N 38°21'35'' E), 1570 m a.s.l., 02.6.2009; 2 female, 2 male larvae, [20.] TURKEY: Artvin Province, Artvin town, Gorgit Plateau Road, Didrelle Creek (41°25'13.65"N 41°55'29.57"E), 922 m a.s.l., 29.7.2016.

Other material examined (not type): 1 male larva, [3.] GEORGIA: Adjara, vicinity of Khino village, Kintrishi Riv. (41°43'01"N 42°02'41"E), 794 m a.s.l., 19.4.2016; 4 female, 3 male larvae, [5.]; 1 juv. larva, [6.] GEORGIA: Adjara, Kokoleti village, Machakhlistskali Riv. (41°29'54.7"N 41°51'26.1"E), 273 m a.s.l., 17.6.2013; 1 female larva, [7.] GEORGIA: Imereti, small stream, road Sairme-Abastumani (41°51'49.0"N 42°47'21.1"E), 1804 m a.s.l., 30.6.2013; 1 male larva, [10.] GEORGIA: Racha-Lechkhumi and Kvemo Svaneti Region, Legvanta village, Sakora Riv. (42°36'32.28"N 43°27'45.18"E), 837 m a.s.l., 25.8.2009; 4 juv. larvae, [11.] GEORGIA: Adjara, app. 5 km E. Goderdzi Pass, left tributary of Dzindzinstskali Riv. (41°38'00.18"N 42°33'24.6"E), 1801 m a.s.l., 28.8.2009; 1 female larva, [14.]; 1 female larva, [15.]; 1 female larva, ARMENIA: Kotayk Province, Geghard, Azat Riv., July 1975; 4 female larvae, [17.] TURKEY: Giresun Province, Dereli-Kümbet Road, Aksu Stream (40°41'01.08"N 38°26'29.71"E), 557 m a.s.l., 1.6.2009; 1 female larva [18.] TURKEY: Giresun Province, Kümbet Plateau, Aksu Stream, (40°32'22''N 38°24'06''E), 1542 m a.s.l., 2.6.2009.