Published February 18, 2021
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Two sea anemones (Cnidaria: Anthozoa: Actiniaria) from the Southern Ocean with evidence of a deep-sea, polar lineage of burrowing sea anemones
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Gusmão, Luciana C., Rodríguez, Estefanía (2021): Two sea anemones (Cnidaria: Anthozoa: Actiniaria) from the Southern Ocean with evidence of a deep-sea, polar lineage of burrowing sea anemones. Zoological Journal of the Linnean Society 193: 1392-1415, DOI: 10.1093/zoolinnean/zlaa176, URL: https://doi.org/10.1093/zoolinnean/zlaa176
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- https://www.checklistbank.org/dataset/3130
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- urn:lsid:zoobank.org:pub:D000BAA9-51C4-46FE-AC9B-BB361B601FA2
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- urn:lsid:plazi.org:pub:D55D9C65FFB9FFCDFF8FC136FFD4B21F
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- http://publication.plazi.org/id/D55D9C65FFB9FFCDFF8FC136FFD4B21F
References
- Andres A. 1881. Prodromus neapolitanae actinarium faunae addito generalis actiniarum bibliographiae catalogo. Mittheilungen aus der Zoologischen Station zu Neapel 2: 305-371.
- Andres A. 1883. Le Attinie. Atti dell'Accademia de Lincei 14: 211-673.
- Appellof A. 1896. Die Actiniengattungen Fenja, Aegir u. Halcampoides, DAN. Bergens Museums Aarbog 1896: 1-16.
- Barnes DKA, De Grave S. 2001. Ecological biogeography of southern polar encrusting faunas. Journal of Biogeography 28: 359-365.
- Bax N, Cairns S. 2014. Stylasteridae (Cnidaria: Hydrozoa). In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the southern ocean. Cambridge: Scientific Committee on Antarctic Research, 107-112.
- Brandt A, Van de Putte A, Griffiths H. 2014. Southern Ocean benthic deep-sea biodiversity and biogeography. In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the Southern ocean. Cambridge: Scientific Committee on Antarctic Research, 233-239.
- Cairns SD, Bayer FM, Fautin DG. 2007. Galatheanthemum profundale (Anthozoa: Actiniaria) in the western Atlantic. Bulletin of Marine Science 80: 191-200.
- Carlgren O. 1893. Studien uber Nordische Actinien. Kongliga Svenska Vetenskaps-akademiens Handlingar 25: 1-148.
- Carlgren O. 1900. Uber Pentactinia californica n. gen. n. sp. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar 57: 1165-1172.
- Carlgren O. 1921. Actiniaria I. Danish Ingolf-Expedition 5: 1-241.
- Carlgren O. 1925. Zur Kenntnis der Hexacorallen. Zoologischer Anzeiger 65: 87-99.
- Carlgren O. 1927. Actiniaria and Zoantharia. In: Odhner T, ed. Further Zoological Results of the Swedish Antarctic Expedition 1901-1903. Stockholm: P. A. Norstedt & Soner, 1-102.
- Carlgren O. 1928. Actiniaria der Deutschen Tiefsee- Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer 'Valdivia' 1898-1899 22: 125-266.
- Carlgren O. 1931. Zur Kenntnis der Actiniaria Abasilaria. Arkiv fur Zoologi 23: 1-48.
- Carlgren O. 1932. Die Ceriantharien , Zoantharien und Actiniarien des arktischen Gebietes. In: Romer F, Schaudinn F, Brauer A,Arndt W, eds. Eine Zusammenstellung der arktischen Tierformen mit besonderer Berucksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nordliche Eismeer im Jahre 1898. Jena: Gustav Fischer, 255-266.
- Carlgren O. 1938. South African Actiniaria and Zoantharia. Kungliga Svenska Vetenskapsakademiens Handlingar 17: 1-148.
- Carlgren O. 1942. Actiniaria Part II. Danish Ingolf-Expedition 5: 1-92.
- Carlgren O. 1949. A survey of the Ptychodactiaria, Corallimorpharia and Actiniaria. Kungliga Svenska Vetenskapsakademiens Handlingar 1: 1-121.
- Carlgren O. 1951. The actinian fauna of the Gulf of California. Proceedings of the United States National Museum 101: 415-449.
- Carlgren O. 1956. Actiniaria from depths exceeding 6000 meters. Galathea Reports 2: 9-16.
- Carlgren O. 1959. Corallimorpharia and Actiniaria with description of a new genus and species from Peru. Reports of the Lund University Chile Expedition 1948-49, 38. Lunds Universitets Arsskrift (NF) Adv 2 56: 1-39.
- Clarke A, Crame JA. 1992. The Southern Ocean benthic fauna and climate change: a historical perspective. Philosophical Transactions of the Royal Society B 338: 299-309.
- Clarke A, Crame J. 2010. Evolutionary dynamics at high latitudes: speciation and extinction in polar marine faunas. Philosophical Transactions of the Royal Society B 365: 3655-3666.
- Clarke A, Johnston NM. 2003. Antarctic marine benthic diversity. Oceanography and Marine Biology 41: 47-114.
- Clubb JA. 1902. Actiniae.With an account of their peculiar brood chambers. In: Report on collections of natural history made in the Antarctic Regions during the voyage of the 'Southern Cross'. London: British Museum (Natural History), 294-309.
- Dall WH. 1903. A new genus of Trochidae. Nautilus 17: 61-62.
- Daly M, Lipscomb DL, Allard MW. 2002. A simple test: evaluating explanations for the relative simplicity of the Edwardsiidae (Cnidaria: Anthozoa). Evolution; international journal of organic evolution 56: 502-510.
- Daly M, Chaudhuri A, Gusmao L, Rodriguez E. 2008. Phylogenetic relationships among sea anemones (Cnidaria: Anthozoa: Actiniaria). Molecular Phylogenetics and Evolution 48: 292-301.
- Daly M, Gusmao LC, Reft AJ, Rodriguez E. 2010. Phylogenetic signal in mitochondrial and nuclear markers in sea anemones (Cnidaria, Actiniaria). Integrative and Comparative Biology 50: 371-388.
- Danielssen DC. 1890. Actinida. Den Norske Nordhavs- Expedition 1876-1878. Zoologi. Christiania: Grondahl & Son.
- Darriba D, Taboada GL, Doallo R, Posada D. 2012. jModelTest 2: more models, new heuristics and parallel computing. Nature Methods 9: 772.
- De Broyer C, Koubbi P. 2014. The biogeography of the Southern Ocean. In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the Southern Ocean. Cambridge: Scientific Committee on Antarctic Research, 2-9.
- Diaz A, Feral JP, David B, Saucede T, Poulin E. 2011. Evolutionary pathways among shallow and deep-sea echinoids of the genus Sterechinus in the Southern Ocean. Deep-Sea Research II 58: 205-211.
- Dunn DG. 1983. Some Antarctic and sub-Antarctic sea anemones (Coelenterata: Ptychodactiaria and Actiniaria). Biology of the Antarctic Seas XIV, Antarctic Research Series 39: 1-67.
- Edmands S, Fautin DG. 1991. Redescription of Aulactinia veratra n. comb. (= Cnidopus veratra) (Coelenterata: Actiniaria) from Australia. Records of the Western Australian Museum (Perth) 15: 59-68.
- Eleaume M, Hemery LG, Roux M, Ameziane N. 2014. Southern Ocean crinoids. In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the Southern Ocean. Cambridge: Scientific Committee on Antarctic Research, 208-212.
- Fautin DG. 1988. The importance of nematocysts to actiniarian taxonomy. In: Hessinger DA, Lenhoff HM, eds. The biology of nematocysts. New York: Academic Press, 487-500.
- Fautin DG. 2016. Catalog to families, genera, and species of orders Actiniaria and Corallimorpharia (Cnidaria: Anthozoa). Zootaxa 4145: 1-449.
- Fautin DG, Barber BR. 1999. Maractis rimicarivora, a new genus and species of sea anemone (Cnidaria: Anthozoa: Actiniaria: Actinostolidae) from an Atlantic hydrothermal vent. Proceedings of the Biological Society of Washington 112: 624-631.
- Fautin DG, Hessler RR. 1989. Marianactis bythios, a new genus and species of actinostolid sea anemone (Coelenterata: Actiniaria) from the Mariana vents. Proceedings of the Biological Society of Washington 102: 815-825.
- Fedorov A, Beichel R, Kalpathy-Cramer J,Finet J,Fillion- Robin JC, Pujol S, Bauer C, Jennings D, Fennessy F, Sonka M, Buatti J, Aylward S, Miller JV, Pieper S, Kikinis R. 2012. 3D Slicer as an image computing platform for the Quantitative Imaging Network. Magnetic Resonance Imaging 30: 1323-1341.
- Geller JB, Walton ED. 2001. Breaking up and getting together: evolution of symbiosis and cloning by fission in sea anemones (Genus Anthopleura). Evolution; International Journal of Organic Evolution 55: 1781-1794.
- Goloboff PA, Farris J, Nixon K. 2008. TNT, a free program for phylogenetic analysis. Cladistics 24: 774-786.
- Gosse PH. 1858. Synopsis of the families, genera, and species of the British Actiniae. Annals and Magazine of Natural History 1: 414-419.
- Gosse PH. 1859. Characters and descriptions of some new British sea-anemones. Annals and Magazine of Natural History 3: 46-50.
- Grajales A, Rodriguez E. 2016. Elucidating the evolutionary relationships of the Aiptasiidae, a widespread cnidariandinoflagellate model system (Cnidaria: Anthozoa: Actiniaria: Metridioidea). Molecular Phylogenetics and Evolution 94: 252-263.
- Griffiths HJ, Barnes DKA, Linse K. 2009. Towards a generalised biogeography of the Southern Ocean benthos. Journal of Biogeography 36: 162-177.
- Guindon S, Gascuel O. 2003. A simple, fast, and accurate algorithm to estimate large phylogenies by maximum likelihood. Systematic Biology 52: 696-704.
- Guindon S, Dufayard JF, Lefort V, Anisimova M, Hordijk W, Gascuel O. 2010. New algorithms and methods to estimate maximum-likelihood phylogenies: assessing the performance of PhyML 3.0. Systematic Biology 59: 307-321.
- Gusmao LC, Daly M. 2010. Evolution of sea anemones (Cnidaria: Actiniaria: Hormathiidae) symbiotic with hermit crabs. Molecular Phylogenetics and Evolution 56: 868-877.
- Gusmao LC, Grajales A, Rodriguez E. 2018. Sea anemones through X-rays: visualization of two species of Diadumene (Cnidaria, Actiniaria) using micro-CT. American Museum Novitates 3907: 1-45.
- Gusmao LC, Berniker K, Van Deusen V, Rodriguez E. 2019. Halcampulactidae (Actiniaria, Actinostoloidea), a new family of burrowing sea anemones with external brooding from Antarctica. Polar Biology 42: 1271-1286.
- Gusmao LC, Van Deusen V, Daly M, Rodriguez E. 2020. Origin and evolution of the symbiosis between sea anemones (Cnidaria, Anthozoa, Actiniaria) and hermit crabs, with additional notes on anemone-gastropod associations. Molecular Phylogenetics and Evolution 148: 106805.
- Gutt J, Hosie G, Stoddart M. 2010. Marine life in the antarctic. In: McIntyre AD, ed. Life in the World's Oceans: Diversity, Distribution, and Abundance. Oxford: Blackwell Publishing Ltd., 203-220.
- Helmuth B, Veit RR, Holberton R. 1994. Long-distance dispersal of a subantarctic brooding bivalve (Gaimardia trapesina) by kelp-rafting. Marine Biology 120: 421-426.
- Hertwig R. 1882. Die actinien der challenger expedition. Jena: Gustav Fischer.
- Janosik AM , Mahon AR , Halanych KM . 2011 . Evolutionary history of Southern Ocean Odontaster sea star species (Odontasteridae; Asteroidea). Polar Biology 34: 575-586.
- Janussen D, Downey RV. 2014. Porifera. In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic Atlas of the Southern Ocean. Cambridge: Scientific Committee on Antarctic Research, 94-102.
- Jennison BL 1981. Reproduction in three species of sea anemones from Key West, Florida. Canadian Journal of Zoology 59: 1708-1719.
- Katoh K, Standley DM. 2013. MAFFT multiple sequence alignment software v.7: improvements in performance and usability. Molecular Biology and Evolution 30: 772-780.
- Katoh K, Rozewicki J, Yamada KD. 2017. MAFFT online service: multiple sequence alignment, interactive sequence choice and visualization. Brief in Bioinformatics 20: 1160-1166.
- Kearse M, Moir R, Wilson A, Stones-Havas S, Cheung M, Sturrock S, Buxton S, Cooper A, Markowitz S, Duran C, Thierer T, Ashton B, Meintjes P, Drummond A. 2012. Geneious Basic: an integrated and extendable desktop software platform for the organization and analysis of sequence data. Bioinformatics (Oxford, England) 28: 1647-1649.
- Krug AZ, Jablonski D, Valentine JW. 2009. Signature of the end-cretaceous mass extinction in the modern biota. Science 323: 767-771.
- Koubbi P, De Broyer C, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, Stoddart M, Swadling KM, Wadley V. 2014. Conclusions: present and future of Southern Ocean biogeography. In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the Southern Ocean. Cambridge: Scientific Committee on Antarctic Research, 470-475.
- Kozlov AM, Darriba D, Flouri T, Morel B, Stamatakis A. 2018. RAxML-NG: a fast scalable, and user-friendly tool for maximum likelihood phylogenetic inference. Bioinformatics 35: 4453-4455.
- Larson P. 2017. Brooding sea anemones (Cnidaria: Anthozoa: Actiniaria): paragons of diversity in mode, morphology, and maternity. Invertebrate Zoology 136: 92-112.
- Larson P, Daly M. 2015. 'Putting names with faces': a description of Epiactis handi sp. nov. helps to solve taxonomic confusion in species of the sea anemone Epiactis (Actiniaria, Actiniidae). Journal of Marine Biological Association of the United Kingdom 95: 913-928.
- Lauretta D, Haussermann V, Brugler MR, Rodriguez E. 2014. Isoparactis fionae sp. n. (Cnidaria: Anthozoa: Actiniaria) from southern Patagonia with a discussion of the family Isanthidae. Organisms Diversity and Evolution 14: 31-42.
- Lefort V, Longueville J, Gascuel O. 2017. SMS: Smart model selection in PhyML. Molecular Biology and Evolution 34: 2421-2424.
- Lopez-Gonzalez PJ, Rodriguez E, Gili J, Segonzac M. 2003. New records on sea anemones (Anthozoa: Actiniaria) from hydrothermal vents and cold seeps. Zoologische Verhandelingen 345: 215-243.
- Lutken C. 1875. A revised catalogue of the Anthozoa and Calycozoa of Greenland. In: Jones TR, ed. Manual of the natural history, geology, and physics of Greenland and the neighbouring regions. London: Her Majesty's Stationery Office, 186-187.
- McMurrich JP. 1893. Report on the Actiniae collected by the United States Fish Commission Steamer Albatross during the winter of 1887-1888. Proceedings of the United States National Museum 16: 119-216.
- Miller MA, Pfeiffer W, Schwartz T. 2010. Creating the CIPRES Science Gateway for inference of large phylogenetic trees. Proceedings of the Gateway Computing Environments Workshop (GCE), New Orleans, 1-8.
- Munilla T, Soler Membrives A. 2009. Check-list of the pycnogonids from Antarctic and sub-Antarctic waters: zoogeographic implications. Antarctic Science 21: 99-111.
- Nikula R, Fraser CI, Spencer HG, Waters JM. 2010. Circumpolar dispersal by rafting in two subantarctic kelpdwelling crustaceans. Marine Ecology Progress Series 405: 221-230.
- Nikula R, Spencer HG, Waters JM. 2013. Passive rafting is a powerful driver of transoceanic gene flow. Biology Letters 9: 20120821.
- Ostman C. 2000. A guideline to nematocyst nomenclature and classification, and some notes on the systematic value of nematocysts. Scientia Marina 64: 31-46.
- Patarnello T, Verde C, di Prisco G, Bargelloni L, Zane L. 2011. How will fish that evolved at constant sub-zero temperatures cope with global warming? Notothenioids as a case study. Bioessays: News and Reviews in Molecular, Cellular and Developmental Biology 33: 260-268.
- Pearse JS, Mooi R, Lockhart SJ, Brandt A. 2009. Brooding and species diversity in the Southern Ocean: selection for brooders or speciation within brooding clades? In: Krupnik I, Lang MA, Miller SE, eds. Smithsonian at the Poles: Contributions to International Polar Year Science. Proceedings of Smithsonian at the Poles Symposium, Smithsonian Institution, Washington, DC, 3-4 May 2007. Washington, DC: Smithsonian Institution Scholarly Press, 181-196.
- Pena Cantero AL. 2014. Benthic hydroids (Cnidaria: Hydrozoa). In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the Southern Ocean. Cambridge: Scientific Committee on Antarctic Research, 103-116.
- Pena Cantero AL, Garcia Carrascosa AM. 1999. Biogeographical distribution of the benthic thecate hydroids collected during the Spanish 'Antartida 8611' expedition and comparison between Antarctic and Magellan benthic hydroid faunas. Scientia Marina 63: 209-218.
- Pfeffer G. 1889. Zur Fauna von Sud-Georgien. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 6: 39-55.
- Presnell JK, Schreibman MP. 1997. Humason's animal tissue techniques. Baltimore: Johns Hopkins University Press.
- Primo C, Vazquez E. 2007. Zoogeography of the Antarctic ascidian fauna in relation to the sub-Antarctic and South America. Antarctic Science 19: 321-336.
- Quattrini AM, Rodriguez E, Faircloth BC, Cowman PF, Brugler MR, Farfan GA, Hellberg ME, Kitahara MV, Morrison CL, Paz-Garcia DA,Reimer JD,McFadden CS. 2020. Palaeoclimate ocean conditions shaped the evolution of corals and their skeletons through deep time. Nature Ecology & Evolution 4: 1531-1538.
- Rafinesque CS. 1815. Analyse de la nature ou tableau de l'univers et des corps organises. Palermo: Rafinesque.
- Riemann-Zurneck K. 1978. Actiniaria des Sudwestatlantik IV. Actinostola crassicornis (Hertwig, 1882) mit einer Diskussion verwandter Arten. Veroffentlichungen des Institutes fur Meeresforschung Bremerhaven 17: 65-85.
- Rintoul S. 2007. Antarctic circumpolar current. In: Riffenburgh B, ed. Encyclopedia of the Antarctic. New York: Routledge, 234-239.
- Rodriguez E, Daly M. 2010. Phylogenetic relationships among deep-sea and chemosynthetic sea anemones: Actinoscyphiidae and Actinostolidae (Actiniaria: Mesomyaria). PLoS One 5: e1095.
- Rodriguez E, Fautin DG. 2014. Antarctic hexacorals (Cnidaria: Anthozoa: Hexacorallia). In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the Southern Ocean. Cambridge: Scientific Committee on Antarctic Research, 113-116.
- Rodriguez E, Lopez-Gonzalez PJ. 2002. A new species of Halcampella (Actiniaria: Halcampoididae) from the eastern Weddell Sea and the Antarctic Peninsula. Scientia Marina 66: 43-52.
- Rodriguez E, Lopez-Gonzalez PJ. 2013. New records of Antarctic and sub-Antarctic sea anemones (Cnidaria, Anthozoa, Actiniaria and Corallimorpharia) from the Weddell Sea, Antarctic Peninsula, and Scotia Arc. Zootaxa 3624: 1-100.
- Rodriguez E,Lopez-Gonzalez PJ,Gili J. 2007. Biogeography of Antarctic sea anemones (Anthozoa, Actiniaria): what do they tell us about the origin of the Antarctic benthic fauna? Deep-Sea Research II 54: 1876-1904.
- Rodriguez E, Castorani CN, Daly M. 2008. Morphological phylogeny of the family of Actinostolidae (Anthozoa: Actiniaria) with description of a new genus and species of hydrothermal vent sea anemone redefining the family Actinoscyphidae. Invertebrate Zoology 22: 439-452.
- Rodriguez E, Lopez-Gonzalez PJ, Daly M. 2009. New family of sea anemones (Actiniaria, Acontiaria) from deep polar seas. Polar Biology 32: 703-717.
- Rodriguez E, Barbeitos M, Daly M, Gusmao LC, Haussermann V. 2012. Toward a natural classification: phylogeny of acontiate sea anemones (Cnidaria, Anthozoa, Actiniaria). Cladistics 1: 1-18.
- Rodriguez E, Orejas C, Lopez-Gonzalez PJ, Gili JM. 2013. Reproduction in the externally brooding sea anemone Epiactis georgiana in the Antarctic Peninsula and the Weddell Sea. Marine Biology 160: 1-14.
- Rodriguez E, Barbeitos MS, Brugler MR, Crowley LM, Grajales A, Gusmao L, Haussermann V, Reft A, Daly M. 2014. Hidden among sea anemones: first comprehensive phylogenetic reconstruction of the order Actiniaria (Cnidaria: Anthozoa: Hexacorallia) reveals a novel group of hexacorals. PLoS One 9: 1-15.
- Sanamyan NP, Sanamyan KE. 2007. Deep-water Actiniaria from East Pacific hydrothermal vents and cold seeps. Invertebrate Zoology 4: 83-102.
- Sanamyan N, Sanamyan K. 2012. Edwardsia sojabio sp. n. (Cnidaria: Anthozoa: Actiniaria: Edwardsiidae), a new abyssal sea anemone from the Sea of Japan. Deep Sea Research II 86-87: 225-230.
- Sanamyan NP, Sanamyan KE, Lundsten L. 2015. Two new deepwater sea anemones (Cnidaria: Anthozoa: Actiniaria) Sicyonis heliodiscus sp. n. (Actinostolidae) and Hormathia pacifica sp. n. (Hormathiidae) from Pacific. Invertebrate Zoology 12: 131-149.
- Sanamyan NP,Sanamyan KE,McDaniel N,Bocharova ES. 2018. First record of two genera of sea anemones (Cnidaria, Actiniaria), Octineon and Edwardsiella from the North Pacific Ocean. Invertebrate Zoology 15: 1-18.
- Saucede T , Pierrat B, Danis B, David B. 2014. Biogeographic processes in the Southern Ocean. In: De Broyer C, Koubbi P, Griffiths HJ, Raymond B, d'Udekem d'Acoz C, Van de Putte A, Danis B, David B, Grant S, Gutt J, Held C, Hosie G, Heuttmann F, Post A, Ropert-Coudert Y, eds. Biogeographic atlas of the Southern Ocean. Cambridge: Scientific Committee on Antarctic Research, 456-463.
- Stephenson TA. 1918. Coelenterata. Part I. - Actiniaria. Natural History Reports on British Antarctic ('Terra Nova') Expedition 1910 5: 1-68.
- Stephenson TA. 1920. On the classification of Actiniaria. Part I.-Forms with acontia and forms with a mesogloeal sphincter. Quarterly Journal of Microscopical Science 64: 425-574.
- Stephenson TA. 1922. On the classification of Actiniaria. Part III. Definitions connected with the forms dealt with in Part II. Quarterly Journal of Microscopical Science 66: 247-319.
- Stephenson TA. 1928. The British sea anemones, Vol. 1. London: Ray Society.
- Studer T. 1879. Ubersicht der Anthozoa Alcyonaria, welche wahrend der Reise S.M.S. Gazelle um die erde gesammelt wurden. Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin 1878: 632-688.
- Van Praet M. 1990. Gametogenesis and the reproductive cycle in the deep-sea anemone Paracalliactis stephensoni (Cnidaria: Actiniaria). Journal of the Marine Biological Association of the United Kingdom 70: 163-172.
- Verrill AE. 1883. Reports on the Anthozoa, and on some additional species dredged by the 'Blake' in 1877-1879, and by the U. S. Fish Commission Steamer 'Fish Hawk' in 1880- 82. Bulletin of the Museum of Comparative Zoology (Harvard University) 11: 1-72.
- Verrill AE. 1899. Descriptions of imperfectly known and new Actinians, with critical notes on other species, IV. American Journal of Science and Arts 7: 205-218.
- Yanagi K, Fujii T, Hirose M. 2015. Redescription of the sea anemone Exocoelactis actinostoloides (Cnidaria: Anthozoa: Actiniaria) based on a topotypic specimen collected from Tokyo Bay, Japan. Species Diversity 20: 199-209.