(Asteraceae), 20.iii.1983 (Martin

#4092) ETYMOLOGY. The chosen species name is keris, a type of Malay dagger with a wavy blade, similar to the rather sinuous apical section of the axial process which issues from each of the large compound pores in this species. The word is used as noun in apposition.

COMMENTS. C. keris has puparia whose large compound pores each possess a very long, sword­like, axial process which extends well beyond the puparial margin, and the puparial outline is usually exaggeratedly asymmetrical. Most puparia develop parallel to, and contiguous with, the midrib or another major leaf vein, the puparia themselves then being asymmetrical, with large compound pores only on the side furthest from the adjacent leaf vein; two specimens from Panamá (on Heterocondylus and Mouriri, see material examined, above) are symmetrical in outline, have most of their large compound pores paired on either side of the body (Fig. 19), and probably each developed away from a major vein. Variation in the number of large compound pores also occurs, with either two or three abdominal pores present (Figs 17, 18): one sample from Nicaragua includes individuals with two and with three abdominal large pores. However, all other characters remain entirely typical, and the number and degree of pairing of the large compound pores is regarded as intraspecific variation – see Table 1. There is considerable size variation amongst the study material, but there is no evidence of puparial sexual dimorphism.

All puparia occur singly, and collection of a few specimens always involves examination of the lower surfaces of many leaves. In this respect, C. keris is similar to several other species of Ceraleurodicus and Nealeurodicus (personal field observations).