Type locality. — Borneo, E Sabah, Danum Valley.

Type material.— Holotype ♂ from the type locality, Danum Valley Field Centre, primary forest, ground trap at foot of buttress tree, 10-15.v. 1991, C.L. Deeleman and M. Goodnight; paratypes: 1 ♂ same data, 1 ♂ in dung trap, Andrew Davis 1991, 1 ♂, 1 ♀, ground trap in forest which was partly logged 12 years earlier and 1 ♂ in primary unlogged forest, same data, 1989-1990, Th. Borghouts.

Other material. — Borneo, E Kalimantan, N of Balikpapan, 1 ♂ “Sepaku”, stand of degraded primary rainforest, leaf litter, 16.vii. 1979, PR. Deeleman; Samboga, N of Balikpapan, 1 ♂, secondary light forest, leaf litter, 6.viii. 1980, P.R. and C.L. Deeleman; SE Kalimantan, Aranio district, 2 ♀, primary forest of Gunung Pisang, 5 km S of Belangiang village, 60 km SE of Banjarmasin, 200-250 m, sifting leaf litter, 11-15.xi.1996, M. Kuntner (SIW). Two males from the type locality from pitfall traps (1989-1990) were sacrificed to making SEM pictures (Platnick, Bonaldo).

Diagnosis.— Very similar to S. borneensis, distinct by the nodule-covered area of carapace which is extended over the entire surface including the centre; nodules are rounded, larger, higher and denser. Differences in the male palp are slight: the lower branch of the tibial apophysis is longer and distal margin partly straight. In females the difference in the vulva is obvious, compare fig. 755 and fig. 751.

Description. — MALE, holotype. Total length 3.70 mm, other males from Danum and other localities 3.60-4.00 mm. Carapace length 1.80 mm, width 1.35 mm, height at coxae II 0.60 mm, head width 0.80 mm, width eye group 0.55 mm; abdomen 1.90 mm long including collar, 1.20 mm wide. Leg lengths: leg I 4.70 mm (1.25-1.75-1.00-0.70), leg II 4.10 mm (1.15-1.45-0.85-0.65), leg III 3.60 mm (1.00-1.15-0.90-0.55), leg IV 5.65 mm (1.50-1.75-1.50-0.90), palp 0.62-0.25-0.50 including apophysis- 0.75 mm. Carapace, sternum and mouthparts dark reddish brown, entirely covered with dense rows of nodules (fig. 753), leg colouring stereotyped as in borneensis. Dorsal scutum uniform chestnut, smooth and shiny, distally dark grey with white spot in front of anal tubercle; abdomen with white underside and a dark circle around spinnerets. Anterior eyes equal and close together, AME 11/ 2 x d ALE; posterior eye row faintly procurved, posterior eyes roughly equidistant; PME not smaller than other eyes. Leg spination: spines rather small compared with other species, femur I with 0-1-0pl, femur IV proximally with a larger dorsal spine, tibia I basically with 6 ventral pairs, tibia II with 5, but frequently loss of spines occurs, uni- or bilaterally; tibia III with 0-2-0v, tibia IV with 1-2-0 or 0-2-0 and and 0-0-1 rl, metatarsi I and II with 2-2-2-2v, metatarsus III with 2-0v and 0-1 rl, metatarsus IV with 2- 2v or 1-2v and 0-1 rl. Dorsal scutum covering almost all of abdomen (scutum incomplete in the male from Sepaku), collar of ventral scutum with three grooves. Palp (figs 737, 754) very similar to that of borneensis, for differences see diagnosis; embolus as long as in borneensis.

FEMALE, paratype. Total length 5.00 mm (females from Gunung Pisang 6.25 and 6.75 mm respectively). Carapace length 2.50 mm, width 1.80 mm, head width 1.10 mm, width eye group 0.55 mm; abdomen 2.80 mm long, 2.10 mm wide, epigyne 0.45 mm wide, 0.45 mm long. Leg lengths: leg I 5.90 mm (1.70-2.20-1.25-0.75), leg II 5.50 mm (1.70-1.90­ 1.10-0.80), leg III 4.60 mm (1.25-1.50-1.10-0.75), leg IV 6.80 mm (1.75-2.25-1.80-1.00), palp 0.75-0.30-0.40- 0.70 mm. Carapace, eyes, mouthparts, sternum and legs as in the ♂; abdomen covered with scutum over 2/ 3 in the middle a pair of round light spots with a central black dot, posterior dorsum dark grey with white pre-anal spot, sides dark grey with white band just behind ventral scutum. Epigyne longer than wide, openings located posterolaterally (fig. 755), vulva fig. 756, spermathecae relatively small and wide apart. In females from Mount Pisang, the spermathecae are a little larger Distribution.— At present only known from Borneo in a zone of forest, primary and degraded, of appr. 100 km stretching all along the eastern coast. See also distribution of borneensis. Etymology. — From Latin bulbus, bulb: bearing bulbs (nodules) on the carapace.