Published July 5, 2021
| Version v1
Journal article
Open
Conservation of Peninsular Malaysia's Karst Herpetofauna: A review of herpetological discoveries, research trends, and challenges
Authors/Creators
Description
Quah, Evan S.H., Grismer, L. Lee, Anuar, M.S. Shahrul (2021): Conservation of Peninsular Malaysia's Karst Herpetofauna: A review of herpetological discoveries, research trends, and challenges. Raffles Bulletin of Zoology 69: 235-252, DOI: 10.26107/RBZ-2021-0019
Files
source.pdf
Files
(6.8 MB)
| Name | Size | Download all |
|---|---|---|
|
md5:1911c612162f5e57745caf2be4c2b07b
|
6.8 MB | Preview Download |
Linked records
Additional details
Identifiers
- LSID
- urn:lsid:zoobank.org:pub:E670B4C8-23AE-4758-BB1F-FFDA105E1892
- LSID
- urn:lsid:plazi.org:pub:1911C612162F5E57745CAF2BFFC2B07B
Related works
- Has part
- Figure: 10.5281/zenodo.5352176 (DOI)
- Figure: 10.5281/zenodo.5352178 (DOI)
- Figure: 10.5281/zenodo.5352180 (DOI)
References
- Ali CA, Ahmad N, Abdullah M, Leman MS, Komoo I & Latiff A (eds.) (2010) Kilim Karst Geoforest Park, Langkawi Geopark, Malaysia, Selangor. Langkawi Research Centre, Institute for Environment and Development (LESTARI), Universiti Kebangsaan Malaysia, UKM Bangi, 122 pp.
- Annandale N, Brown JC & Gravely FH (1913) The limestone caves of Burma and the Malay Peninsula. Journal of the Asiatic Society of Bengal, 9(10): 391-423.
- Berry PY (1970) The food of the giant toad Bufo asper. Zoological Journal of the Linnean Society, 49(2): 61-68.
- Berry PY (1975) The amphibian fauna of Peninsular Malaysia. Tropical Press, Kuala Lumpur, 127 pp.
- Boulenger GA (1900) Description of a new lizard from the Batu Caves, Selangor. Journal of the Bombay Natural History Society, 13: 335.
- Boulenger GA (1903) Report on the Batrachians and Reptiles. In: Annandale N & Robinson HC (eds.) Fasciculi Malayensis, anthropological and zoological results of an expedition to Perak and the Siamese Malay States, 1901-1902, undertaken by Nelson Annandale and Herbert C Robinson. Zoology, Part I. Williams and Norgate, London, pp. 131-176.
- Boulenger GA (1912) A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Reptilia and Batrachia. Taylor & Francis, London, 294 pp.
- Bristowe WS (1952) The Arachnid Fauna of the Batu Caves in Malaya. Annals and Magazine of Natural History, 12(5): 697-706.
- Brooks S (2002) Thailand 1998-1999. Thungyai Naresuan W.L. Sanctuary & Thung Salaeng Luang N.P. The International Caver, 2001: 74-76.
- Bullock JA (1963) Notes on the cave faunas of two limestone massifs in the Taman Negara. Malayan Nature Journal, 17: 46-52.
- Bullock JA (1971) Fauna of Gua Anak Takun. Malayan Nature Journal, 24(1970-71): 95-97.
- Bullock JA (1973) Cave Biology in Malaysia. Malayan Nature Journal, 25(1971-72): 135-141.
- Cabauatan JG, Ramos MT, Taggueg JB, Callueng AM & Tumaliuan SS (2014) Assessment of faunal diversity on selected caves of the Northern Sierra Madre Natural Park (NSMNP), Northern Cagayan Valley, Philippines. International Journal of Agricultural Technology, 10(3): 631-649.
- Chakravaty R & Isaac S (2013) Beobachtungen an Gonyosoma oxycephalum (Boie, 1827) beim Erbeuten von Hohlenfledertieren auf den Andamanen, Indien. Sauria, 36(2): 55-58.
- Chan KO, Belabut D & Ahmad N (2010) A revised checklist of the amphibians of Peninsular Malaysia. Russian Journal of Herpetology, 17: 202-206.
- Chan KO & Grismer LL (2008) A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Selangor, Peninsular Malaysia. Zootaxa, 1877: 49-57.
- Chan KO, Grismer LL, Sharma DS, Belabut D & Ahmad N (2009) New herpetofaunal records for Perlis State Park and adjacent areas. Malayan Nature Journal, 61(4): 277-284.
- Charlton T (2020) A Guide to Snakes of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 299 pp.
- Chen L, Kangning X & Guangmei W (2013) Process of biodiversity research of karst areas in China. Acta Ecologica Sinica, 33(4): 192-200.
- Chung KF, Leong WC, Rubite RR, Repin R, Kiew R, Liu Y & Peng C-I (2014) Phylogenetic analyses of Begonia sect. Coelocentrum and allied limestone species of China shed light on the evolution of Sino-Vietnamese karst flora. Botanical Studies, 55: 1-15.
- Clements R, Sodhi NS, Schilthuizen M & Ng PKL (2006) Limestone karsts of Southeast Asia: Imperilled arks of biodiversity. BioScience, 56: 733-742.
- Clements R, Ng PKL, Lu XX, Ambu S, Schilthuizen M & Bradshaw CJA (2008) Using biogeographical patterns of endemic land snails to improve conservation planning of limestone karsts. Biological Conservation, 141: 2751-2764.
- Davis HR, Bauer AM, Jackman TR, Nashriq I & Das I (2019) Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia. Zootaxa, 4614: 331-352.
- Davis HR, Grimser LL, Cobos AJ, Murdoch ML, Sumarli AX, Anuar S, Muin MA, Wood Jr PL & Quah ESH (2018) Checklist of the Herpetofauna of Hutan Lipur Gunung Senyum, Pahang, Peninsular Malaysia. Russian Journal of Herpetology, 25(3): 207-220.
- Day MJ & Urich PB (2000) An assessment of protected karst landscapes in Southeast Asia. Cave and Karst Science, 27: 61-70.
- De Alban JDT, Leong BPI, Venegas-Li R, Connette GM, Jamaludin J, Latt KT, Oswald P, Reeder C & Webb EL (2021) Conservation beyond the existing protected area network is required to improve species and habitat representation in a global biodiversity hotspot. Biological Conservation, 257: 109105.
- De Rooij N (1915) The Reptiles of the Indo-Australian Archipelago. I: Lacertilia, Chelonia, Emydosaura. E. L. Brill, Leiden, 384 pp.
- Dehling JM (2011) A new karst-dwelling species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia. Zootaxa, 2737: 49-60.
- Dover C (ed.) (1929) Fauna of the Batu Caves, Selangor. Journal of the Federated Malay States Museums, 14: 325-387.
- Doyle ME (1969) Factors affecting distribution of fauna in Gua Pondok. Malayan Nature Journal, 23: 21-26.
- Dring JC (1979) Amphibians and reptiles from northern Trengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum Natural History (Zoology), 34: 181-241.
- Dubois A, Ohler A & Pyron RA (2021) New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians (Lissamphibia). Megataxa, 5(1): 1-738.
- Dunn FL (1965) Gua Anak Takun: Ecological observations. Malayan Nature Journal, 19: 75-87.
- Ellis M (2020) The fauna of the limestone caves of Thailand to August 2020. Phetchabun, Thailand, 128 pp.
- Ellis M & Pauwels OS (2012) The bent-toed geckos (Cyrtodactylus) of the caves and karst of Thailand. Cave and Karst Science, 39(1): 16-22.
- Frost DR (2021) Amphibian Species of the World: an Online Reference. Version 6.1. Electronic Database. American Museum of Natural History, New York, USA. Available from: https:// amphibiansoftheworld.amnh.org/index.php (Accessed 21 February 2021).
- Geyer E, Schmidt F & Jeutter PW (eds.) (2005) Speleological Expedition Gunung Lanno. Verein fur Hohlenkunde in Obersteier, Austria, 240 pp.
- Gillieson D (2005) Karst in Southeast Asia. In: Gupta A (ed.) The Physical Geography of Southeast Asia. Oxford University Press, Oxford, pp. 157-176.
- Goddard A & Ellis M (2018) SMCC Expedition to Phitsanulok and Phetchabun 2018. Shepton Mallet Caving Club Journal, 13(8): 337-360.
- Gray A (2001) Thailand and Malaysia Expedition 26 December 2000 to 14 January 2001. Axbridge Caving Group Journal, March 2001: 9-64.
- Grismer LL (2011a) Lizards of Peninsular Malaysia, Singapore and Their Adjacent Archipelagos. Chimaira, Frankfurt am Main, 728 pp.
- Grismer LL (2011b) Field Guide to the Amphibians and Reptiles of the Seribuat Archipelago, Peninsular Malaysia. Chimaira, Frankfurt am Main, 239 pp.
- Grismer LL, Belabut DM, Quah SH, Chan KO, Wood Jr PL & Hashim R (2014a) A new species of karst forest adapted Benttoed Gecko (Genus Cyrtodactylus) belonging to the C. sworderi complex from a threatened karst forest in Perak, Peninsular Malaysia. Zootaxa, 3755(5): 434-466.
- Grismer LL, Chan KO, Anuar S & Muin MA (2014b) Cyrts in the city: a new Bent-toed Gecko (Genus Cyrtodactylus) is the only endemic species of vertebrate from Batu Caves, Selangor, Peninsular Malaysia. Zootaxa, 3774(4): 381-394.
- Grismer LL, Chan KO, Nurolhuda N & Sumontha M (2008a) A new species of karst dwelling gecko (genus Cnemaspis Strauch 1887) from the border region of Thailand and Peninsular Malaysia. Zootaxa, 1875: 51-68.
- Grismer LL, Grismer JL, Wood Jr PL & Chan KO (2008b) The distribution, taxonomy, and redescription of the geckos Cnemaspis affinis (Stoliczka 1887) and C. flavolineata (Nicholls 1949) with descriptions of a new montane species and two new lowland, karst-dwelling species from Peninsular Malaysia. Zootaxa, 1931: 1-24.
- Grismer LL, Norhayati A, Chan KO, Belabut D, Muin MA, Wood Jr PL & Grismer JL (2009) Two new diminutive species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa, 2019: 40-56.
- Grismer LL & Quah ESH (2019) An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa, 4545(2): 230-248.
- Grismer LL, Quah ESH, Anuar S, Muin MA, Wood Jr PL & Nor SAM (2014c) A diminutive new species of cave dwelling Wolf Snake (Colubridae: Lycodon Boie, 1826) from Peninsular Malaysia. Zootaxa, 3815(1): 51-67.
- Grismer LL, Wood Jr PL, Anuar S, Davis HR, Cobos AJ & Murdoch ML (2016a) A new species of karst forest Bent-toed Gecko (genus Cyrtodactylus Gray) not yet threatened by foreign cement companies and a summary of Peninsular Malaysia's endemic karst forest herpetofauna and the need for its conservation. Zootaxa, 4061(1): 1-17.
- Grismer LL, Wood Jr PL, Anuar S, Grismer MS, Quah ESH, Murdoch ML, Muin MA, Davis HR, Aguilar C, Klabacka R, Cobos AJ, Aowphol A & Sites Jr JW (2016b) Two new Bent-toed Geckos of the Cyrtodactylus pulchellus complex from Peninsular Malaysia and multiple instances of convergent adaptation to limestone forest ecosystems. Zootaxa, 4105(5): 401-429.
- Grismer LL, Wood Jr PL, Anuar S, Quah ESH, Muin MA, Mohamed M, Chan KO, Sumarli AX, Loredo AI & Heinz HM (2014d) The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia. Zootaxa, 3786(3): 359-381.
- Grismer LL, Wood Jr PL, Anuar S, Riyanto A, Norhayati A, Muin MA, Sumontha M, Grismer JL, Chan KO, Quah ESH & Pauwels OSG (2014e) Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa, 3880: 1-147.
- Grismer LL, Wood Jr PL, Aowphol A, Cota M, Grismer MS, Murdoch M, Aguilar C & Grismer JL (2016c) Out of Borneo, again and again: biogeography of the Stream Toad genus Ansonia Stoliczka (Anura: Bufonidae) and the discovery of the first limestone cave-dwelling species. Biological Journal of the Linnean Society, 120: 371-395.
- Grismer LL, Wood Jr PL, Le MD, Quah ESH & Grismer JG (2020a) Evolution of habitat preference in 243 species of Bent-toed Geckos (Genus Cyrtodactylus Gray, 1827) with a discussion of karst habitat conservation. Ecology and Evolution, 10: 13717-13730.
- Grismer LL, Wood Jr PL, Mohamed M, Chan KO, Heinz HM, Sumarli, AS-I, Chan JA & Loredo AI (2013) A new species of karst-adapted Cnemaspis Strauch, 1887 (Squamata: Gekkonidae from a threatened karst region in Pahang, Peninsular Malaysia. Zootaxa, 3746(3): 463-472.
- Grismer LL, Wood Jr PL, Poyarkov NA, Le MD, Karunarathna S, Chomdej S, Suwannapoom C, Qi S, Liu S, Che J, Quah ESH, Kraus F, Oliver PM, Riyanto A, Pauwels OSH & Grismer JL (2021a) Karstic Landscapes are Foci of Species Diversity in the World's Third Largest Vertebrate Genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata; Gekkonidae). Diversity, 13(5): 183.
- Grismer LL, Wood Jr PL, Poyarkov NA, Le MD, Kraus F, Agarwal I, Oliver PM, Nguyen SN, Nguyen TQ, Karunarathna S, Welton LJ, Stuart BL, Luu VQ, Bauer AM, O'Connell KA, Quah ESH, Chan KO, Ziegler T, Ngo H, Nazarov RA, Aowphol A, Chomdej S, Suwannapoom C, Siler CD, Anuar S, Tri NV & Grismer JL (2021b) Phylogenetic partitioning of the thirdlargest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology, 71: 101-154.
- Grismer LL, Wood Jr PL, Quah ESH, Anuar S, Muin MA, Sumontha M, Norhayati A, Bauer AM, Wangkulangkul S, Grismer JL & Pauwels OSG (2012) A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa, 3520: 1-55.
- Grismer LL, Wood Jr PL, Quah ESH, Grismer MS, Thura MK, Oaks JR & Lin A (2020b) Two new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from a karstic archipelago in the Salween Basin of southern Myanmar (Burma). Zootaxa, 4718(2): 151-183.
- Grismer LL, Wood Jr PL, Quah ESH & Thura MK (2020c) Origin, diversity, and conservation of karst-associated Bent-toed Geckos (Genus Cyrtodactylus) in Myanmar (Burma). Israel Journal of Ecology & Evolution, 2020: 1-7.
- Grismer LL, Wood Jr PL, Quah ESH, Thura MK, Oaks JR & Lin A (2020d) Four new Burmese species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from distantly related parapatric clades from the Shan Plateau and Salween Basin. Zootaxa, 4758(1): 45-82.
- Grismer LL, Wood Jr PL, Quah ESH, Thura MK, Oaks JR & Lin A (2019) A new species of Bent-toed Gecko (Squamata, Gekkonidae, Cyrtodactylus) from the Shan Plateau in eastern Myanmar (Burma). Zootaxa, 4624(3): 301-321.
- Grismer LL, Wood Jr PL, Thura MK, Quah ESH, Grismer MS, Murdoch ML, Espinoza RE & Aung L (2018a) A new Cyrtodactylus Gray, 1827 (Squamata, Gekkonidae) from the Shan Hills and the biogeography of Bent-toed Geckos from eastern Myanmar. Zootaxa, 4446(4): 477-500.
- Grismer LL, Wood Jr PL, Thura MK, Quah ESH, Murdoch ML, Grismer MS, Herr MW, Lin A & Kyaw H (2018b) Three more new species of Cyrtodactylus (Squamata: Gekkonidae) from the Salween Basin of eastern Myanmar underscore the urgent need for the conservation of karst habitats. Journal of Natural History, 52(19-20): 1243-1294.
- Grismer LL, Wood Jr PL, Thura MK, Zin T, Quah ESH, Murdoch ML, Grismer MS, Lin A, Kyaw H & Lwin N (2018c) Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from limestone habitats in east-central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism. Zoological Journal of the Linnean Society, 182: 862-959.
- Grismer LL, Wood Jr PL, Thura MK, Zin T, Quah ESH, Murdoch ML, Grismer MS, Lin A, Kyaw H & Lwin N (2017) Phylogenetic taxonomy of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) with descriptions of three new species from Myanmar. Journal of Natural History, 52(13-16): 881-915.
- Grismer LL, Wood Jr PL, Zug GR, Thura MK, Grismer MS, Murdoch ML, Quah ESH & Lin A (2018d) Two more new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from the Shan Hills of eastern Myanmar (Burma). Zootaxa, 4483(2): 295-316.
- Grismer LL, Youmans TM, Wood Jr PL, Ponce A, Wright SB, Jones BS, Johnson R, Sanders KL, Gower DJ, Norsham SY & Lim KKP (2006) Checklist of the herpetofauna of Pulau Langkawi, Malaysia, with comments on taxonomy. Hamadryad, 30(1&2): 61-74.
- Hartmann L, Mecke S, Kieckbusch M, Mader F & Kaiser H (2016) A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Muller, 1895). Zootaxa, 4067(5): 552-568.
- Hymeir K (1998) The Survey of Limestone Caves of Perlis with Management Recommendations. Part 1: Caves of the Perlis State Park. Project No. MY 0067. WWF Malaysia, Petaling Jaya, 90 pp.
- Inger RF, Stuart BL & Iskandar DT (2009) Systematics of a widespread Southeast Asian frog, Rana chalconota (Amphibia: Anura: Ranidae). Zoological Journal of the Linnean Society, 155: 123-147.
- Inger RF, Stuebing RB, Grafe TU & Dehling JM (2017) A Field Guide to the Frogs of Borneo. Third Edition. Natural History Publications (Borneo), Kota Kinabalu, 228 pp.
- IUCN (2021) The IUCN Red List of Threatened Species. Version 2020-3. https://www.iucnredlist.org (Accessed 03 February 2021).
- Kiew BH (1979) A Note on the Frogs and Toads of Batu Caves. Malayan Nature Journal, 33(1): 67-69.
- Kiew BH (1985) A brief report on the cave fauna of the eight limestone massifs between Sungai Kenyam and Sungai Trenggan, Taman Negara. In Gua Kepayang '84. Taman Negara Cave expedition. Malaysian Naturalist, 38(3): 29.
- Kiew R (1991) The limestone flora. In: Kiew R (ed.) The State of Nature Conservation in Malaysia. Malaysian Nature Society, Kuala Lumpur, pp. 42-50.
- Kiew R (2014) Checklist of vascular plants from Batu Caves, Selangor, Malaysia. Check List, 10(6): 1420-1429.
- Komericki A, Aung L, Ngwe L & Momberg F (2020) Myanmar karst key biodiversity areas. Fauna and Flora International Special Report, 104 pp.
- Latinne A, Waengsothorn S, Herbreteau V & Michaux JR (2011) Thai limestone karsts: An impending biodiversity crisis. In: Proceedings of the 1st Environment Asia International Conference on Environmental Supporting in Food and Energy Security: Crisis and Opportunity. Thai Society of Higher Education Institutes on Environment, Bangkok, Thailand, pp. 176-187.
- Liew TS, Price L & Clements GR (2016) Using Google Earth to improve the management of threatened limestone karst ecosystems in Peninsular Malaysia. Tropical Conservation Science, 9(2): 903-920.
- Luu VQ, Bonkowski M, Nguyen TQ, Le MD, Calame T & Ziegler T (2018) A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos. Revue suisse de Zoologie, 125(2): 263-276.
- Luu VQ, Bonkowski M, Nguyen TQ, Le MD, Schneider N, Ngo HT & Ziegler T (2016) Evolution in karst massifs, cryptic diversity among bent-toed geckos along the Truong Son Range with descriptions of three new species and one new country record from Laos. Zootaxa, 4107: 101-140.
- Manchi S & Sankaran R (2009) Predators of swiftlets and their nests in the Andaman and Nicobar Islands. Indian Birds, 5: 118-120.
- Mane AM & Manchi SS (2018) Civets in the Limestone Caves of Andaman Islands: Notes on the Distribution and Natural History. Ambient Science, 5(1): 6-10.
- Manthey U & Denzer W (2014) Sudostasiatische Anuren im Fokus: Spezies der Gattung Microhyla Tschudi, 1838 (Anura: Microhylidae: Microhylinae). Sauria, 36(3): 3-21.
- Manthey U, Denzer W & Quah ESH (2016) Sudostasiatische Anuren im Fokus: Microhyla palmipes Boulenger, 1897 und Microhyla superciliaris Parker, 1928 der Malaiischen Halbinsel. Sauria, 38: 23-30.
- McClure HE (1965) Microcosms of Batu Caves. Malayan Nature Journal (Anniversary Issue), 19: 65-74.
- McClure HE, Lim BL & Winn SE (1967) Fauna of the Dark Cave, Batu Caves, Kuala Lumpur, Malaysia. Pacific Insects, 3: 399-428.
- Mecke S, Kieckbusch M, Hartmann L & Kaiser H (2016) Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa, 4175(4): 343-365.
- Monroe WH (1970) A Glossary of Karst Terminology, Washington: US Geological Survey Water Supply Paper 1899-K. GV Print, USA, 29 pp.
- Moseley M, Lim TW & Lim TT (2012) Fauna reported from Batu caves, Selangor, Malaysia: annotated checklist and bibliography. Cave and Karst Science, 39: 77-92.
- Nazarov RA, Pauwels OSG, Konstantinov EL, Chulisov AS, Orlov NL & Poyarkov NA (2018) A new karst-dwelling bent-toed gecko (Squamata, Gekkonidae, Cyrtodactylus) from Xiangkhoang Province, northeastern Laos. Zoological Research, 38: 1-16.
- Nuneza OM, Calizo-Enguito MR, Labajo-Villantes Y & Ponce AG (2015) Species richness and endemism of cave herpetofauna in Northern Mindanao, Philippines. ELBA Bioflux, 7(1): 10-24.
- Oakley P & Smart D (2002) Reconnaissance cave expedition in the Thungyai Naresuan West Wildlife Sanctuary, Thailand April 2002. Royal Forest Department, Bangkok, 12 pp.
- Orlov NL, Ryabov SA, Thanh BN & Cuc HT (2004) A new species of Trimeresurus (Ophidia: Viperidae: Crotalinae) from karst region in Central Vietnam. Russian Journal of Herpetology, 11(2): 139-149.
- Panitvong N, Lauhachinda V, Saithong S & Temchai T (2012) Ecology of Cyrtodactylus sumonthai Bauer, Pauwels & Chanhome, 2002 (Reptilia: Squamata: Gekkonidae): A karst dwelling bent-toed gecko from South-eastern Thailand. Raffles Bulletin of Zoology, 60: 569-582.
- Parker HW (1928) The brevicipitid frogs of the genus Microhyla. Annals and Magazine of Natural History, 10(2): 473-499.
- Paton JR (1961) A brief account of the geology of the limestone hills of Malaya. Bulletin of the Raffles Museum, 26: 66-75.
- Pauwels OS, David P, Chimsunchart C & Thirakhupt K (2003) Reptiles of Phetchaburi Province, Western Thailand: a list of species, with natural history notes, and a discussion on the biogeography at the Isthmus of Kra. Natural History Journal of Chulalongkorn University, 3(1): 23-53.
- Pauwels OSG, Sumontha M, Panitvong N & Varaguttanonda V (2014) Cyrtodactylus khelangensis, a new cavedwelling benttoed gecko (Squamata: Gekkonidae) from Lampang Province, northern Thailand. Zootaxa, 3755: 584-594.
- Poyarkov NA, Van Nguyen T, Van Duong T, Gorin VA & Yang JH (2018) A new limestone-dwelling species of Micryletta (Amphibia: Anura: Microhylidae) from northern Vietnam. PeerJ, 6: e5771.
- Price L (1992) The Batu Caves, Kuala Lumpur, Malaysia. International Caver, 3: 24-28.
- Price L (1998) Kenong Rimba. Malaysian Naturalist, 51(3&4): 28-31.
- Price L (2001) Caves and karst of Peninsular Malaysia: A register. Gua Publications, Kuala Lumpur, 98 pp.
- Price L (2014a) Caves and Karst of Peninsular Malaysia. A Register. Second Edition. Berliner Hohlenkundliche Berichte, 54: 1-43.
- Price L (2014b) Species Diversity and Food-web Complexity in the Caves of Malaysia. Ambient Science, 1(2): 1-8.
- Price L & Li Y-W (2008) The Cave Racer, Orthiophris taeniurus. Cave and Karst Science, 34(3): 129-134.
- Quah ESH, Grismer LL, Wood Jr PL & Shahrul AMS (2019) The discovery and description of a new species of Bent-toed Gecko of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa, 4668(1): 51-75.
- Quah ESH, Wood Jr PL, Shahrul AMS, Muin MA & Grismer LL (2020) A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia with an updated checklist of the herpetofauna of Tuba Island. Zootaxa, 4767(1): 138-160.
- Ridley HN (1898) The white snake of the Selangor Caves. Journal of the Straits Branch of the Royal Asiatic Society, 31: 99-101.
- Ridley HN (1899) Report: Appendix to Caves of the Malay Peninsula.-Report of the Committee consisting of Sir W. H. Flower (Chairman), Mr. H. N. Ridley (Secretary), Dr. R. Hanitsch, Mr. Clement Reid, and Mr. A Russell Wallace, appointed to explore certain caves in the Malay Peninsula, and to collect their living and extinct Fauna. Report of the Sixty- Eighth Meeting of the British Association for the Advancement of Science held at Bristol in September 1898. John Murray, London, pp. 572-582.
- Riyanto A, Bauer AM & Yudha DS (2014) A new small karstdwelling species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Java, Indonesia. Zootaxa, 3785(4): 589-599.
- Schilthuizen M, Liew TS, Elahan BB & Lackman Ancrenaz I (2005) Effects of karst forest degradation on pulmonate and prosobranch land snail communities in Sabah, Malaysian Borneo. Conservation Biology, 19(3): 949-954.
- Siler CD, Alcala AC, Diesmos AC & Brown RM (2009) A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from Eastern Samar Island, Philippines. Herpetologica, 65(1): 92-104.
- Siler CD, Diesmos AC, Linkem CW, Diesmos MLL & Brown RM (2010) A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from central Luzon Island, Philippines. Zootaxa, 2482: 49-63.
- Smart D (2002) Thungyai Naresuan cave survey project: Final report. Royal Forest Department, Bangkok, 180 pp.
- Smart D & Cunningham R (2002) Caves of Southern Thailand. Unpublished report. Royal Forest Department, Bangkok, 33 pp.
- Smith MA (1930) The reptilia and amphibia of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. A supplement to G. A. Boulenger's Reptilia and Batrachia, 1912. Bulletin of the Raffles Museum, 3: 1-149.
- Smith RJ (1995) Exercise Tham Farang Thailand 1992 Report. Combined Services Caving Association, United Kingdom, 49 pp.
- Steiner H (2002) Fauna list - Thailand/Malaysia 2000/2001. The International Caver, 2001: 72-73.
- Struebig MJ, Kingston T, Zubaid A, LeComber SC, Mohd-Adnan A, Turner A & Rossiter SJ (2009) Conservation importance of limestone karst outcrops for Palaeotropical bats in a fragmented landscape. Biological Conservation, 142: 2089-2096.
- Stuart BL, Inger RF & Voris HK (2006) High levels of cryptic species diversity revealed by sympatric lineages of Southeast Asian forest frogs. Biology Letters, 2: 470-474.
- Suwannapoom C, Sumontha M, Tunprasert J, Ruangsuwan T, Pawangkhanant P, Korost DV & Poyarkov NA (2018) A striking new genus and species of cave-dwelling frog (Amphibia: Anura: Microhylidae: Asterophryinae) from Thailand. PeerJ, 6: e4422.
- Tolentino PJS, Navidad JRL, Angeles MD, Fernandez DAP, Villanueva ELC, Obena RDR & Buot Jr IE (2020) Biodiversity of forests over limestone in Southeast Asia with emphasis on the Philippines. Biodiversitas Journal of Biological Diversity, 21(4): 1597-1613.
- Uetz P, Freed P & Hosek J (2021) The Reptile Database. Available from: http://www.reptile-database.org (Accessed 01 February 2021).
- Vermeulen JJ (1994) Notes on the non-marine molluscs of the island of Borneo 6. The genus Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae), part 2. Basteria, 58: 75-191.
- Vermeulen J & Whitten T (1999) Biodiversity and Cultural Property in the Management of Limestone Resources-Lessons from East Asia. World Bank, Washington D.C., 120 pp.
- Vogel G, Nguyen TQ, Kingsada P & Ziegler T (2012) A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8: 344-352.
- Wood Jr PL, Grismer LL, Muin MA, Anuar S, Oaks JR & Sites Jr JW (2020) A new potentially endangered limestone-associated Bent-toed Gecko of the Cyrtodactylus pulchellus (Squamata: Gekkonidae) complex from northern Peninsular Malaysia. Zootaxa, 4751(3): 437-460.
- Wood Jr PL, Quah ESH, Anuar S & Muin MA (2013) A new species of lowland karst dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from northwestern Peninsular Malaysia. Zootaxa, 3691(5): 538-558.
- Yong HS (1960) Field Biology: (d) Fauna of the Dark Cave (Batu Caves). Scientific Victorian, 1960: 42-46.
- Yussof S (1997) The natural and other histories of Batu Caves. Malaysian Nature Society, Kuala Lumpur, 55 pp.
- Ziegler T, Hendrix R, Vu NT, Vogt M, Forster B & Dang NK (2007) The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key. Zootaxa, 1493(21): 1-40.
- Ziegler T, Ohler A, Vu NT, Le KQ, Nguyen XT, Dinh HT & Bui NT (2006) Review of the amphibian and reptile diversity of Phong Nha - Ke Bang National Park and adjacent areas, central Truong Son, Vietnam. In: Vences M, Kohler J, Ziegler T & Bohme (eds.) Herpetologia Bonnensis II. Proceedings of the 13th Congress of the Societas Europaea Herpetologica. Societas Europaea Herpetologica, Germany, pp. 247-262.
- Zimmerer J (2004) Nature Guide, Langkawi. Sakti Mega Enterprise, Langkawi, 184 pp.